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Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
Great white sharks (Carcharodon carcharias) males beneath the surface. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Bigfin reef squid (Sepioteuthis lessoniana) capturing a pelagic shrimp with long red antennae, in midwater, at night. Anilao, Batangas, Luzon, Philippines
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
RF - Green sea turtle (Chelonia mydas) beneath clouds. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
A digital composite of a Pygmy Seahorse (Hippocampus bargibanti) swimming over a seafan (Muricella sp.). Pygmy seahorses are small, most less than 15mm in total length
An Australian Sea Lion (Neophoca cinerea) bathing in the sun at the surface. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Giant Pacific Manta Ray (Manta birostris) portrait with Remora fish (Echeneididae) Roca Partida, Revillagigedos Islands, Mexico. East Pacific Ocean. February
Draughtsboard Shark / Australian Swellshark (Cephaloscyllium laticeps) swims through a giant kelp forest (Macrocystis pyrifera)
A large Great hammerhead shark (Sphyrna mokarran), about 4m in length, cruises overhead. Bahamas, Tropical West Atlantic Ocean
Purple short tentacle mushroom coral (Fungia sp. ) close up of polyps. Lembeh StraitPurple short tentacle mushroom coral (Fungia sp.) close up of polyps. Lembeh Strait, North Sulawesi, Indonesia
Pink anemonefish (Amphiprion perideraion) in a purple Magnificent sea anemone (Heteractis magnifica). Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Red boring sponge (Cliona delitrix) three excurrent apertures covered in Sponge zoanthids (Parazoanthus parasiticus) East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Gorgonian / Fan coral (Solenocaulon akalyx), close up, West Papua, Indonesia
Yellow cluster anemone (Parazoanthus axinellae) surrounded by Encrusting sponge (Spirastrella cunctatrix) San Pietro Island, Sardinia, Italy, Mediterranean, July
Starfish close up detail, Blairgowrie, Mornington Penisular, Victoria, Australia. Port Philip Bay, Bass Strait
Close-up of the polyps out of a coral tree (Tubastraea coccinea) at night, Baja California peninsula, Mexico. Sea of Cortez
Orange cup coral (Tubastrea coccinea) Malpelo Island National Park, UNESCO World Heritage Site, Colombia, East Pacific Ocean
Coral clam (Pedum spondyloideum) close up, Maldives, Indian Ocean
Yellow tube sponge (Aplysina fistularis) on a coral reef, with the camera rotated during the long exposure. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
A tiny amphipod (Iphimedia obesa) living on Dead mans fingers soft coral (Alcyonium digitatum) Loch Carron, Ross and Cromarty, Scotland, UK, April
Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines
Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean
Emperor angelfish (Pomacanthus imperator) swimming past soft corals (Dendronephthya sp]), Red Sea, Egypt. January
Flatworm (Maiazoon orsaki) Raja Ampat, Irian Jaya, West Papua, Indonesia, Pacific Ocean
Light Bulb Sea Squirt (Clavelina lepadiformis). Channel Islands, UK, June
A Large / Pot Bellied Seahorse (Hippocampus abdominalis) perched in nets. Manly, Sydney, New South Wales, Australia, March
RF- Boulder star coral (Montastrea annularis) showing fluorescent green coloration, photographed under blue light at night, to excite fluorescent pigments in the coral, East End, Grand Cayman
Brain coral (Diploria strigosa) photographed under blue light to excite the fluorescence, Little Cayman, Cayman Islands, British West Indies, Caribbean Sea
DELETE Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Southern stingray (Dasyatis americana) swimming over sand in shallow water, split level photo with blue sky and clouds. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
School of Blackfin barracuda (Sphyraena qenie), Kimbe Bay, West New Britain, Papua New Guinea
Yellow and blueback fusilier (Caesio teres) schooling over coral reef, West Papua, Indonesia
RF - Young lionfish (Pterois volitans) swimming near surface hunting silversides, at dusk. Beacon Rock, Sha ab Mahmood, Sinai, Egypt. Red Sea
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Sunbeams below the surface of the sea, Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Yellow and blueback fusilier (Caesio teres) shoal, Similan Islands, Andaman Sea, Thailand
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Bigeye trevally or jack (Caranx sexfasciatus) shoal, Sipadan, Malaysia
Blue sea star (Linckia laevigata) on coral reef, Fiji, South Pacific
Atlantic bluefin tuna (Thunnus thynnus) within tuna farm, containing around 1000 per net. Saint Pauls Bay, Malta. Mediterranean Sea
Close up of pattern of adult Emperor angelfish (Pomacanthus imperator) Tulamben, Bali, Indonesia. Java Sea
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water, Ras Katy, Sharm El Sheikh, Sinai, Egypt. Gulf of Aqaba, Red Sea