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Ascidian (Rhopalaea sp) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Goby (Bryaninops sp) on coral branch, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Reef and shoal of Jewel fairy basslet (Pseudanthias squamipinnis) South Point dive site, Sanganeb reef, Sudan, Red Sea
Underwater view of African tigerfish (Hydrocynus vittatus), predatory fish in the Okavango river with Water lily (Nymphaea nouchali var. caerulea) and other aquatic plants
Nile crocodile (Crocodylus niloticus) on the river bed, Okavango River, Okavango Delta, Botswana. Photographed for The Freshwater Project
Underwater view of Brown bear (Ursus arctos) paw fishing for Sockeye salmon (Oncorhynchus nerka) with paw outstretched. Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August
Split level view of water in cave in Wadi Al Shab, Al Sharqiyah South Governorate, Sultanate of Oman. February. Photographed for The Freshwater Project
Underwater view of Sungai Melinau River, Gunung Mulu National Park, Sarawak, Borneo, Malaysia. Photographed for The Freshwater Project
Rangimairewhenua or Blue Lake, sacred to the Maori, Nelson Lakes National Park, South Island, New Zealand, February 2013. Photographed for The Freshwater Project
Head and gills of juvenile Scalloped hammerhead shark (Sphyrna Lewini), Kane ohe Bay, Hawaii, Pacific Ocean
Skeleton shrimp (Caprella septentrionalis) mating Novaya Zemlya, Russian Arctic, July
Galapagos sea lions (Zalophus wollebaeki) hunting tuna. A group of the sea lion bulls have learnt to herd Pelagic yellowfin tuna into a small cove, trapping them
Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp)Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp.) diving towards coral reef to avoid predators. North Ari Atoll, Maldives. Indian Ocean
Oriental sweetlips (Plectorhinchus vittatus) pair. South Ari Atoll, Madlives. Indian Ocean
Reticulate stargazer (Uranoscopus sp. ) portrait. Ambon Bay, Ambon, Maluku Archipelago, IndonesiaReticulate stargazer (Uranoscopus sp.) portrait. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Hawksbill turtle (Eretmochelys imbricata) pair circling each other above coral reef. Seven Mile Beach, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Scalloped hammerhead shark (Sphyrna lewini) Hawaii
Purple stinger (Pelagia noctiluca), Hawaii
Sea cucumber (Bohadschia argus) which has ejected a part of its internal organs called Cuvierian tubules. These significantly sticky strings are a defense mechanism designed to deter predators
Bobbit worm (Eunice sp. ) emerges from its hole in black sand at night to feed. BitungBobbit worm (Eunice sp.) emerges from its hole in black sand at night to feed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Spotfin Burrfish (Chilomycterus reticulatus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Pearlfish (Carapus acus) outside of host Sea cucumber (Stichopus regalis) at night. These species have a commensal relationship, with the fish living in the gut of the sea cucumber. Catalonia, Spain
Common fangtooth (Anoplogaster cornuta) deep sea species from Atlantic Ocean close to Cape Verde. Captive
Spaghetti garden eels (Gorgasia maculata) stretching up out of their burrows on a rubble slope, with Mysiid shrimp swimming around their bodies near the seabed. Dumaguete, Dauin, Negros, Philippines
Bluestreak cleaner wrasse (Labroides dimidiatus) just swimming out of mouth of Oblique / Diagonal banded sweetlips (Plectorhinchus lineatus) whilst cleaning / feeding, New Caledonia. Pacific Ocean
Tassled scorpionfish (Scorpaenopsis oxycephala) swimming just above sea bed, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Lanternfish (Lepidophanes guentheri) - deepsea species showing bioluminescence
Alligator snapping turtle (Macroclemys temmincki) fishing by luring a fish with the vermiform, worm-like, tongue, USA, captive
A male Leafy Seadragon (Phycodurus eques) swimming. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
Pfeffers Flamboyant Cuttlefish (Metasepia pfefferi) West Papua, Indonesia, April 2007
Hawaiian pom-pom / boxer crab (Lybia edmondsoni)with anemones (Triactis sp) that it carries around holding with the claws and using them for defense, waving in front of the possible aggressor
Giant ostracod {Gigantocypris mulleri} from between 718 and 1864m, Mid-Atlantic Ridge, North Atlantic Ocean
Giant ostracod {Gigantocypris mulleri} from between 867 and 1805m, Mid-Atlantic Ridge, North Atlantic Ocean
Close up of mouth sucker of Japanese / Arctic Lamprey {Lethenteron japonicum} Japan
Atlantic wolffish {Anarrhichus lupus} Sognefjord, Norway
Shoal of Lake Ohrid bleak (Alburnus scoranza) swimming amongst Giant reeds (Arundo donax) along the shore of Lake Ohrid, Lagadin region, Galicica National Park, Macedonia, June 2009
Male Monk seal (Monachus monachus) Deserta Grande, Desertas Islands, Madeira, Portugal, August 2009, Critically endangered species
Jellyfish, Mull, Scotland, June 2009
Ornate wrasse (Thalassoma pavo) shoal, Deserta Grande, Desertas Islands, Madeira, Portugal, August 2009
Basking shark (Cetorhinus maximus) feeding just below the surface, Mull, Scotland, June 2009
One of the deeper artificial reefs, Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009
Caramote prawn (Penaeus kerathurus) Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009
Common octopus (Octopus vulgaris) in hole, Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009
Wide-eyed flounder (Bothus podas) on seabed, Marine Reserve, Monaco, Mediterranean Sea, July 2009
Scorpionfish (Scorpaena porcus) lying on the artificial reef, Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009
Shoal of Mediterranean / Blue damselfish (Chromis chromis) Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009
White water lily (Nymphea alba) in flower, viewed from below the surface, Lake Skadar, Lake Skadar National Park, Montengro, May 2008
Fragrant water lilies (Nymphaea odorata) in Lake Skadar, Montenegro, May 2008
