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Yellow / spiny seahorse (Hippocampus guttulatus) female sheltering is meadow of common Eelgrass (Zostera marina) Studland Bay, Dorset, UK, August
Bed of Common mussels (Mytilus edulis) growing on rocks, feeding, Megavissey, Cornwall, UK, May
Rear view of Basking shark (Cetorhinus maximus) feeding on plankton in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June
Basking shark (Cetorhinus maximus) disappears back into the blue in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June
Common guillemot (Uria aalge) swimming underwater, Farne Islands, Northumberland, UK, July
White nudibranch (Eubranchus farrani) on kelp frond, Lunna Kirk, Shetland Islands, Scotland, UK, June
A forest of Sea cucumbers (Psolus phantapus) feeding, extended up out of the sediment in Scottish sea loch, Loch Duich, Ross and Cromarty, Scotland, UK, April
Lesser spotted catshark / Dogfish shark (Scyliorhinus canicula) hiding amongst seaweeds, Babbacombe Bay, Devon, UK, April
Masked crab (Corystes cassivelaunus) on sandy seabed, Studland Bay, Dorset, UK, May
Compass jellyfish (Chrysaora hysoscella) swimming over a rocky reef, Plymouth, Devon, UK, August
Leachs spider crab (Inachus phalangium) sheltering from the current beneath tentacles of Snakelocks anemone (Anemonia viridis) Selsey, West Sussex, UK, May
Brown Trout (Salmo trutta) in disused quarry, Jackdaw Quarry, Capernwray, Carnforth, Lancashire, UK, August
Grey seal (Halichoerus grypus) swimming amongst kelp, Farne Islands, Northumberland, England, UK, August. 2020VISION Book Plate
Jewel anemone (Corynactis viridis), North Uist, Outer Hebrides, Scotland, UK, May. Small repro only
Great white shark (Carcharodon carcharias) Guadalupe Island or Isla Guadalupe, Pacific Ocean, Mexico, September. Vulnerable species
Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
Bull sharks (Carcharhinus leucas) in Beqa (Benga) Lagoon, a reserve where sharks are fed as part of a commercial diving operation. Viti Levu, Fiji, South Pacific
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Field of Giant plumrose anemones (Metridium giganteum) with a Snakelock anemone / Crimson sea anemone (Cribrinopsis fernaldi) in foreground, Alaska, USA, Gulf of Alaska. Pacific ocean
Detail of the pectoral fin of Roundhead parrotfish (Scarus viridifucatus) coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
School of Bluespine unicornfish (Naso unicornis) gathered during the summer breeding season. Yolanda Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Sperm whale (Physeter macrocephalus) tail fluke above water during dive, Kaikoura, New Zealand, July, Vulnerable species
Golf-ball sized Painted frogfish (Antennarius pictus) waiting to ambush prey disguised as an orange sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Close up portrait of female Lionfish (Pterois volitans) in the late afternoon, when Lionfish are more active. The Alternatives, Sha ab Mahmood, Sinai, Egypt. Red Sea
Male Yellow boxfish (Ostracion cubicus) displaying, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Blue sharks (Prionace glauca) Faial Island, Azores, Portugal, Atlantic Ocean, September
Common dolphins (Delphinus delphis) pair in mating ritual, Atlantic ocean, Portugal, July
School of young European perch (Perca fluviatilis) in Altausseer lake, Austria, July
Bowhead Whale (Balaena mysticetus) rubbing off flaking skin on the ocean bottom, east coast Baffin island, Qikiqtarjuaq, Nunavut, Canada, May
The narrow Nes Canyon, a fault filled with fresh water in the rift valley between the Eurasian and American tectonic plates in northern Iceland
Great Blue shark (Prionace glauca) viewed from just below the surface, Pico Island, Azores, Portugal, Atlantic Ocean
Underwater view of tree, with sun rays entering the water, Fibreno Lake nature reserve, Lazio, Italy
Whale shark (Rhincodon typus) mouth open filter feeding at the surface, Quintana Roo, Yucatan Peninsular, Mexico, Caribbean Sea
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Freshwater Dorado (Salminus brasiliensis) Recanto Ecologico, Rio da Prata, Bonito, Mato Grosso do Sul, Brazil
Manta ray (Manta birostris), San Benedicto, Revillagigedo (Socorro) Islands, Mexico, East Pacific Ocean
Plumose anemones (Metridium senile) and common starfish (Asterias rubens) beneath the sun in a Norwegian Fjord. Gulen, Bergen, Norway. North East Atlantic Ocean
A signal blenny (Emblemaria hypacanthus) displays its showy dorsal fin as it extends out of its tube home. La Paz, Mexico. Sea Of Cortez, East Pacific Ocean
Lemon sole (Microstomus kitt) portrait. Gardur, south west Iceland. North Atlantic Ocean
Scenic view of a bull kelp forest (Nereocystis luetkeana) with sunlight shining through the fronds. Browning Pass, Port Hardy, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
Crack of the continental drift between the North American and Eurasian Plates at Thingvellir, Thingvellir National Park, UNESCO World Heritage Site, Iceland. September 2009
Gannet (Sula bassana) in water, Shetland, UK. July
Bigfin squid (Sepioteuthis lessoniana) hovering in mid water at night. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Slender sapsucking slug (Thuridilla gracilis) on surface of Royal seasquirt (Polycarpa aurata) Aljui Bay, Waigeo Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
