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Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
British World War II BSA M20 motorbike inside Hold 2 of wreck of HMS Thistlegorm, home to pair of Red Sea soldierfish (Myripristis murdjan) and nudibranch (Nembrotha megalocera)
Detail of the filter feeding apendange of Peacock worm (Sabella pavonina) Loch Carron, Ross and Cromarty, Scotland, UK, April
Yellow bellied snapping turtle (Elseya flaviventralis) about to surface for air, Arnhem Escarpment Creek, Northern Territory, Australia. June
Worrells turtle (Emydura subglobosa worrelli), large adult female resting at night, Lawn Hill Creek, Queensland, Australia. January
Sandstone long-necked turtle (Chelodina burrungandjii) actively foraging at night, Pentecost River, Western Australia. April
Spotted eagle ray (Aetobatus narinari), Kicker Rock, San Cristobal Island, Galapagos
Tail of Blue whale (Balaenoptera musculus brevicauda). This may be the pygmy sub-species of blue whale, Balaenoptera musculus. Mirissa, Sri Lanka, Indian Ocean
Lemon shark juvenile (Negaprion brevirostris) trying to feed on the leaves of a red mangrove (Rhizophora mangle). As pups, the sharks have to learn to hunt for themselves and sometimes make mistakes
Double exposure of a Tompot blenny (Parablennius gattorugine) with underwater sun beams. Swanage, Dorset, England, United Kingdom
A reef scene with an orange elephant ear sponge (Ianthella basta) and red sea whip (Ellisella sp) with schooling silversides fish above. Pelee Island, Misool, Raja Ampat, West Papua, Indonesia
Split level photo of mangrove scenery, with hard corals ( including Goniopora sp.; Heliopora sp; Porites sp.) growing below Red mangrove tree: (Rhizophora mangle)
Giant barrel sponge (Xestospongia testudinaria) releases a large cloud of gametes as it spawns on a coral reef, while a Threadfin hawkfish (Cirrhitichthys aprinus) looks
Black grouper (Mycteroperca bonaci) yawning on a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
RF - Open brain coral (Trachyphyllia geoffroyi) lives in symbiosis with zooxanthellae which are responsible for nutrition and the vibrant colours. Lembeh Strait, North Sulaweisi, Indonesia
Big scale sand smelt {Atherina boyeri} captive Delta del Ebro NP, Spain
Coral polyps, Honduras
Large group of Sunflower sea stars (Asterias / Pycnopodia helianthoides) covering rock, pacific coast, Canada, August
Inser Weiher in autumn, a pond with red coloured water, caused by dissolved tannins. Berner Seeland, Berne, Switzerland, November. Photographed for The Freshwater Project
Yellow water-lily (Nuphar lutea) underwater. Lake Morat, Vaud, Switzerland, September. Photographed for The Freshwater Project
Pacific white-sided dolphins (Lagenorhynchus obliquidens) underwater, Queen Charlotte Strait, British Columbia, Canada. September
Giant Pacific octopus (Enteroctopus dofleini) showing arms and suckers, Vernon Rock, Queen Charlotte Strait, British Columbia, Canada. September
A Giant rock scallop (Crassadoma gigantea) encrusted with and surrounded by several other invertebrates animals, including White anemones (Metridium sp.), White mushroom ascidians (Distaplia sp.)
Gooseneck barnacles (Pollicipes Polymerus) amongst kelp, Nakwakto Rapids, Slingsby Channel, British Columbia, Canada. September
Purple and Ochre sea stars (Pisaster ochraceus) preying on Pacific blue mussels (Mytilus trossulus). Also seen are Green Surf Anemones (Anthopleura xanthogrammica)
Sockeye salmon (Oncorhynchus nerka) males fighting in river Adams River, British Columbia, Canada. October
Bull kelp canopy (Nereocystis luetkeana) floating on the surface providing protective habitat for juvenile Rockfishes (Sebastes spp.) Browning Pass, Queen Charlotte Strait, British Columbia, Canada
Wolf eel (Anarrhichthys ocellatus) Croker Rock, Queen Charlotte Strait, British Columbia, Canada. September
Stubby squid (Rossia pacifica) Nigei Island, Queen Charlotte Strait, British Columbia, Canada
Opalescent nudibranchs (Hermissenda crassicornis) and Mushroom compound ascidians (Distaplia occidentalis). Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
Hooded nudibranchs (Melibe leonina) with oral hood for catching planktonic prey. In the Pacific Northwest they are often found clinging to bull kelp (Nereocystis luetkeana)
Sea butterfly (Cymbulia peronii) originally classed as Pseudothecosomata. Cymbuliidae are pelagic sea snails, marine gastropod molluscs. Atlantic Ocean, close to Cape Verde. Controlled conditions
Endemic sponge (Lubomirskia baicalensis) encrusting rocks and branches on the lake floor, Lake Baikal, Siberia, Russia. July
Amphipod (Acanthogammarus victorii) on lake bed, with some swimming, Lake Baikal, Siberia, Russia. Endemic to Lake Baikal
Young Baikal seal (Pusa sibirica) at breathing hole Lake Baikal, Siberia, Russia. April
Baikal seal (Pusa sibirica) underwater, endemic to Lake Baikal, Siberia, Russia. July
Short-snouted seahorse (Hippocampus hippocampus)Tenerife, Canary Islands
Red-lipped blenny (Ophioblennius atlanticus) portrait, Tenerife, Canary Islands
Wide-eyed flounder (Bothus podas) portrait showing eyes above head, Tenerife, Canary Islands
Duckbill ray (Pteromylaeus bovinus) Tenerife, Canary Islands
Tiger moray (Enchelycore anatina) portrait, Tenerife, Canary Islands
Red starfish (Echinaster sepositus) group on sea floor, Tenerife, Canary Islands
Blue starfish (Coscinasterias tenuispina) close up of arm with tube feet, Tenerife, Canary Islands
Atlantic feather star (Antedon bifida) Tenerife, Canary Islands
Cuttlefish (Sepia officinalis) changing its colour to a bright blue, Tenerife, Canary Islands
Green sea turtle (Chelonia mydas) Tenerife, Canary Islands
Short-beaked common dolphin (Delphinus delphis) feeding on Snipe fish (Macroramphosus scolopax) Tenerife, Canary Islands
Sea snails (Smaragdia viridis) on Little neptune seagrass (Cymodocea nodosa) Tenerife, Canary Islands