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A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
RF - Close up detail of the eye and skin pattern of a Giant hawkfish (Cirrhitus rivulatus). Socorro Island, Pacific Ocean
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
RF - Lemon shark (Negaprion brevirostris) swimming through sun rays in the late afternoon. Little Bahama Bank, Bahamas. West Atlantic Ocean
A Leafy Seadragon (Phycodurus eques), photographed from below. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia. Australia
Male Great white shark (Carcharodon carcharias) with sunrays, Guadalupe Island, Mexico, Pacific Ocean
Grooved brain coral (Diploria labyrinthiformis) at night with polyps extended to feed, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
RF - Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia
Detail of Plumose anemone (Metridium senile) in its retracted state at slack water when it is not feeding. Plymouth Sound, Devon, England, UK, English Channel, July
Oman anemonefish / clownfish (Amphiprion omanensis) in a a Bulb-tentacle sea anemone (Entacmaea quadricolor), coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Detail of the gas bladders and fronds of Giant kelp (Macrocystis pyrifera) plant. Santa Barbara Island, the Channel Islands, California, United States of America. East Pacific Ocean
Citron (lemon coral) goby (Gobiodon citrinus) Red Sea
Two Common butterfly fish (Chaetodon chrysurus) Red Sea, Egypt
Great hammerhead shark (Sphyrna mokarran) in shallow water. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Marine iguana (Amblyrhynchus cristatus) underwater. Fernandina Island. Galapagos, Endemic Species
Purple sea pen (Virgularia gustaviana) on sandy sea bed. Rinca, Indonesia
Devonshire cup coral (Caryophyllia smithii) photographed under deep blue light to reveal its fluorescent pigments, showing up as green in this photo. Plymouth Sound, Devon, UK, English Channel, July
Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
Blacktip reef shark (Carcharhinus melanopterus) juvenile in shallow water, Maldives, Indian Ocean
Patters of sunlight on a shallow sandy seabed. North Sound, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Bottlenose dolphins (Tursiops truncatus) approaching with curiosity, Dolphin Reef, Eilat, Israel. Red Sea
Florida Red-bellied Turtle (Pseudemys nelsoni) Rainbow Springs, Rainbow Springs State Park, Florida, USA, March. Taken for the Freshwater Project
Luminescent jellyfish / Mauve stinger (Pelagia noctiluca) in open water, Gozo Island, Malta. Mediterranean Sea
Water chestnut (Trapa natans) viewed from underwater, in Danube Delta, Romania, June
Ruffe (Gymnocephalus cernua) shoal, Danube Delta, Romania, June
European white waterlilies (Nymphaea alba) viewed from underwater in a tributary of Danube Delta, Romania, June
Philippines chromis (Chromis scotochiloptera) swimming in front of reef wall with sponges and featherstars, Komodo National Park, Indonesia
Mediterranean Monk Seal (Monachus monachus) large male of 2.5 m length. Localy identified by Parque Natural da Madeira as " Metade"
Sargassum fish (Histrio histrio) near the surface on floating Sargassum seaweed, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Portuguese Man-of-War (Physalia physalis) close up of tentacles, Sargasso Sea, Bermuda
Morelets crocodile / Central American crocodile (Crocodylus moreletii) hiding in the plants at the surface of the cenote, Car Wash Cenote/ Aktun Ha, Yucatan peninsula, Mexico
Detail of rhinopores and the colour pattern of a toxic Nudibranch (Goniobranchus coi) on a coral reef. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Suwanee freshwater turtle (Pseudemys concinna suwanniensis) swimming beneath trees in a freshwater spring. Devil Spring, Ginnie Springs, High Springs, Florida, USA
Silky shark (Carcharhinus falciformis), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January. Vulnerable
RF - California sea lion (Zalophus californianus) with mouth open close to the camera. The milk teeth of this pup show decay, which is quite usual. Los Islotes, La Paz, Baja California Sur, Mexico
California sea lion (Zalophus californianus) uses a Panamic cushion star (Pentaceraster cumingi) as a toy. The sealions pick up the starfish and then drop them and chase after them as they sink
Southern stingray (Dasyatis americana) female swimming over a shallow sand bank with ripples. Stingray City Sandbar, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Spotted eagle ray (Aetobatus narinari) school, Kagi kandhu, ; North Male Atoll, Maldives, Indian Ocean
Blue spotted trevally (Caranx bucculentus) hunting small fish in the reef, South Ari Atoll, Maldives, Indian Ocean
