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Thread-tail dartgoby (Ptereleotris hanae) Sharp Island or Kiu Tsui Chau, Kiu Tsui Country Park, Hong Kong, China
Northern crested newt (Triturus cristatus) male underwater in a pond, during the mating season. Isere, Cremieu, France
School of Perch (Perca fluviatilis) amongst Water lilies (Nuphar lutea) lit by the suns rays, Lake Bourget, Alps, Savoie, France, June
RF - Soft coral (Dendronephthya sp. ) growing on sea bed. West Papua, IndonesiaRF - Soft coral (Dendronephthya sp.) growing on sea bed. West Papua, Indonesia
Paddle-flap scorpionfish (Rhinopias eschmeyeri) Puerto Galera, Philippines
Devil scorpionfish (Scorpaenopsis diabolus) close up of eye, Pak Lap Tsai, Sai Kung, Hong Kong, China
Stellar rockskipper (Entomacrodus stellifer) hiding in a barnacle. Sai Kung East Country Park, Hong Kong, China
Ocean sunfish (Mola mola) off Halifax, Nova Scotia, Canada. July
Lions mane jellyfish (Cyanea capillata) swept in current over a bed of eel grass, along the Eastern shore of Nova Scotia, Canada, July
Atlantic wolfish (Anarhichas lupus) looking out hole, Bay of Fundy, New Brunswick, Canada, July
Atlantic salmon (Salmo salar) portrait, migrating to spawn in river, Gaspe Peninsula, Quebec, Canada, October
Red water caused by tannins in peat bog flowing out of the Etang de la Gruere, peat bog, Rhine river catchment, Canton of Jura, Switzerland. November 2013. Photographed for the Freshwater Project
Necklace seastar (Fromia monilis) on Sea cucumber (Bohadaschia argus) Yap, Micronesia
Broadclub cuttlefish (Sepia latimanus) in translucent egg casing on the reef, Philippines, Asia
Three spot damselfish (Dascyllus trimaculatus) with Sea anemone home, Yap, Micronesia
Common anemonefish (Amphiprion perideraion) with eggs in Magnificent sea anemone (Heteractis magnifica) Yap, Micronesia
Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) sticking its head into the mouth of a Red Sea fusilier (Caesio lunaris) during a cleaning session. Small Crack, Sha ab Mahmud, Sinai, Egypt. Red Sea
Southern stingray (Dasyatis americana) large female foraging over seagrass in shallow water. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Coral Rabbitfish (Siganus corallinus) portrait, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Cichlids (Cichlidae) in Lake Malawi, Malawi, November. Photographed for The Freshwater Project
Aquatic plants in a spring of the Gacka River, Croatia. October 2012. Photographed for The Freshwater Project
Flower tree soft corals (Umbellulifera sp. ), Triton Bay, near Kaimana, West PapuaFlower tree soft corals (Umbellulifera sp.), Triton Bay, near Kaimana, West Papua, Indonesia
Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluespine unicornfish (Naso unicornis) Shark Reef to Jolande, Ras Mohammed National Park, Egypt, Red Sea
Trumpetfish (Aulostomus maculatus) using a Spanish Hogfish (Bodianus rufus) as a stalking horse. Spanish Hogfish is eating an arm of a starfish. Bonaire, Leeward Antilles, Caribbean
Blue shark (Prionace glauca) off Halifax, Nova Scotia, Canada. July
Veined octopus (Amphioctopus marginatus) between clam shell halves. Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Blue whale (Balaenoptera musculus) diving beneath the surface. Indian Ocean, Sri Lanka
Tawny nurse shark (Nebrius ferrugineus) group, individuals circling each other at dusk. Vavuu Atoll, Maldives. Indian Ocean
Brydes whale (Balaenoptera edeni) Trincomalee, Eastern Province, Sri Lanka, Bay of Bengal, Indian Ocean
Pearlfish (Carapus acus) in anus of Sea cucumber (Stichopus regalis). These species have a commensal relationship, with the fish living in the gut of the sea cucumber. Catalonia, Spain
Sea butterfly (Peraclis bispinosa) captive deep sea species from Atlantic Ocean off Cape Verde
Glass octopus (Vitreledonella richardi). deep sea species from Atlantic Ocean off Cape Verde. Captive
Fangtooth moray eel (Enchelycore anatina) with mouth open, Grand Canaria, Canary Islands, Spain. East Atlantic Ocean
Humboldt squid (Dosidicus gigas) cannibalising another squid of the same species at night off Loreto, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Marbled electric ray (Torpedo marmorata) Bouley Bay, Jersey, British Channel Islands
Sand tiger shark (Carcharias taurus). Caribbean
Aggregation of thousands of Spider crabs (Leptomithrax gaimardii) for moulting, South Australia Basin, Australia. Pacific Ocean
A male Leafy Seadragon (Phycodurus eques) carries a new batch of eggs on his tail. Wool Bay Jetty, Edithburgh, Yorke Peninsula, South Australia, November
Dwarf Minke whale (Balaenoptera acutorostrata) Great Barrier Reef, Australia. Coral Sea
Group of California sea lions (Zalophus californianus) in a kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
RF- Florida manatee (Trichechus manatus latirostrus) portrait in Three Sisters Spring. Crystal River, Florida, USA. February 2010
Yellow water lilies (Nuphar lutea) viewed from underwater, Lake Skadar, Lake Skadar National Park, Montenegro, May 2008
Ratfish / Ghost shark (Chimaera monstrosa) on fjord bed, Trondheimsfjorden, Norway, November 2006
Female Atlantic salmon (Salmo salar) River Orkla, Norway, October 2008
Three Killer whales / Orcas (Orcinus orca) underwater, Kristiansund, Nordmore, Norway, February 2009. WWE INDOOR EXHIBITION
