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Catbells and the edge of Derwentwater bathed in early morning light and autumnal colours, Latrigg, Keswick, The Lake District, Cumbria, UK. November 2016
RF- Beech (Fagus) and Bluebell (Hyacinthoides non-scripta) woodland at Lanhydrock, Cornwall. UK
Polar bear (Ursus maritimus) female coming out the den and shaking off snow. Wapusk National Park, Churchill, Manitoba, Canada, March
Speckled Wood (Pararge aegeria) male flying in habitat, Finland, April
Landscape from Neko Harbour, Andvord Bay. Antarctic Peninsula, Antarctica
RF - Red fox (Vulpes vulpes) in dappled light on snow, Yellowstone National Park, USA. February
Iguasu Falls by moonlight, on the Iguasu River, Brazil / Argentina border. Photographed from the Brazilian side of the Falls. State of Parana, Brazil
Cloud formations lit by low sun. Brechin, Angus, Scotland, December 2012
Cornflowers (Centaurea cyanus) and poppies (Papaver rhoeas) growing on fallow fields near Orvieto, Umbria, Italy. June 2010
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues
RF - Silky shark (Carcharhinus falciformis) swimming beneath the surface of the ocean, with evening sun burst. Gardens of the Queen National Park, Cuba
Grey seal (Halichoerus grypus) young seal silhouetted against the sun as it dives after breathing, Lundy Island, Devon, UK, Bristol Channel, August
Underwater landscape in peat pond, Fribourg, Switzerland, August
RF - Titan triggerfish (Balistoides viridescens) swimming over hard coral (Acropora sp.) gardens. Kri Island, Raja Ampat, West Papua, Indonesia. Dampier Strait, tropical west Pacific Ocean
Broadclub cuttlefish (Sepia latimanus) Komodo National Park, Indonesia
Waikoropupu Springs, Golden Bay, South Island, New Zealand
Amazon river dolphin (Inia geoffrensis), Rio Negro, Amazonia, Brazil
Red Fox (Vulpes vulpes) in wide landscape with Kronotsky Volcano on the horizon. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Mute swan (Cygnus olor) stretching its wings backlit at dawn, Poynton, Cheshire, UK, December
Tsaparang (sometimes known as the mythical Shangri-la) was the capital of the ancient kingdom of Guge in the Garuda Valley, Ngari Prefecture, Western Tibet
Polar bear (Ursus maritimus) adult walking over newly formed pack ice during fall freeze up, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska, USA
A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea
Sunset on Everest (8848m), Sagarmatha National Park (World Heritage UNESCO). Khumbu / Everest Region, Nepal, Himalaya, October 2011
Liberty Bell Mountain at Washington Pass in the North Cascades, in winter, Okanogan National Forest, Washington, USA, November 2011
Grey Heron (Ardea cinerea) taking a fish from water. Kiskunsagi National Park, Hungary, May
Torres del Paine rock towers, Torres del Paine National Park, Patagonia, Chile
Downham Hill, a jurassic limestone outlier on the edge of the Cotswolds AONB conservation area, Gloucestershire, UK, October 2011
Two Least Chipmunks (Tamias minimus) sitting on tree stump feeding, Yellowstone National Park, Wyoming, USA, October, crop of 1373882
A split-level image of a mountain stream, in autumn, with water droplets on lens, Nant Ffrancon Valley, Snowdonia NP, Gwynedd, Wales, UK, October
Hanuman Langur / Northern Plains Grey Langur (Semnopithecus entellus) mother with baby in tree. Kanha National Park, Madhya Pradesh, India
Manta Ray (Manta Birostris) and Devil Ray (Mobula tarapacana) in background, accompanied by Pilot Fish (Naucrates ductor) and Remora. Santa Maria Island, Azores, September
Blue Shark (Prionace glauca) at the sea surface. Santa Maria, Azores, August
Flock of Cape petrels (Daption capense capense) in flight against the sea. Against the sun. South Shetland Islands, Antarctica, January
Lower peaks of the Mont Cervin / Monte Cervino / The Matterhorn massif, Pennine Alps, Switzerland / France border, April 2011
Caribbean reef shark (Carcharhinus perezi) Jardines de la Reina National Park, Cuba
Jersey tiger moth (Euplagia quadripunctaria) large numbers flying to and resting on Sweetgum trees (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011
Bridled nailtail wallaby (Onychogalea fraenata) Queensland, Australia, Endangered species, October
Marbled White Butterfly (Melanargia galathea) resting on grass stem, Devon, UK July. Winner of Seasons category of the BWPA 2011 competition
Pyramid of sea ice catching the first rays of the returning sun in the high Arctic, Svalbard, Spitsbergen, Norway, February 2009. Taken on location for the BBC series, Frozen Planet
Beagle hound in wild purple Violets, Arcadia, Wisconsin, USA
Arctic tern (Sterna paradisaea) adult hovering directly in front of the sun, defending nest from perceived threat, Farne Islands, UK. June Non-ex
A view of a forested inlet lit by low sunlight. Aggro Beach, west coast of Vancouver Island, Canada, September 2010
The snow at Horsetail Falls begins to melt in mid-February, enabling the falls to resume once again. The clear sky and low sun angle ignite the rock wall during sundown
Two Brown bears (Ursus arctos) fighting in woodland wetlands, Kuhmo, Finland, July
Dramatic light on Benijo Beach. Anaga Peninsula, Tenerife, Canary Islands, February 2011
A snowy mountain peak lit by low crepuscular light before sunset. Ordesa National Park, Pyrenees, Aragon, Spain, February 2009
Iceberg against a blue sky. Ilulisat, West Greenland, summer
Apollo butterfly (Parnassius apollo) on thistle head in alpine meadow. Nordtirol, Austrian Alps
