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Deciduous forest with sunlight shining through trees, Monmouthshire, Wales, UK. September
Leopard houndshark (Triakis semifasciata) group swimming amongst rays of light. San Diego, California, USA
Four-spotted chaser dragonfly (Libellula quadrimaculata) covered in dew and backlit by the morning sun, Devon, UK. June
Lemon shark (Negaprion brevirostris) hunting over Turtlegrass (Thalassia testudinum) seagrass bed, view from below. Florida Keys, Florida, USA
Langur monkey (Semnopithecus) climbing tree in misty forest at dawn, India, November
Oceanic whitetip shark (Carcharhinus longimanus) swims in open waters, close to the surface with sun beams. Elphinstone reef, Marsa Alam, Egypt. Red Sea
Young female Grey seal (Halichoerus grypus) beneath sunburst. Farne Islands, Northumberland, UK
Chacma baboon (Papio ursinus) sitting in rainforest in morning light. Jao Reserve, Botswana
Sunbeams play through the roots of red mangrove trees (Rhizophora sp.), encrusted with marine life and home to schoolmaster snappers (Lutjanus apodus)
RF - American crocodile (Crocodylus acutus) swimming through sunrays, above a bed of seagrass ( Thalassia testudinum). Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
RF - Wandering albatross (Diomedea exulans) flying over the rugged coast of South Georgia. South Atlantic. (digitally stitched image)
Kilchurn Castle, sunrise, early morning sunlight, a ruin on a rocky peninsula, the northeastern end of Loch Awe, in Argyll and Bute, Scotland, UK. October
Early morning view, looking southeast from Gordon Channel. Gordon Channel, Queen Charlotte Strait, British Columbia, Canada. September
Sperm whale (Physeter macrocephalus) mother and calf, Sri Lanka. March
Meadow of Neptune seagrass (Posidonia oceanica), Chersonissos, Heraklion, Crete, Greece
Neptune seagrass (Posidonia oceanica) meadow, Samaria National Park, Chania, Crete, Greece
Potato grouper (Epinephelus tukala) in Passe Dubois / Dubois channel, Aldabra, Indian Ocean
Autumn crocus or meadow saffron (Colchicum autumnalis) in Lower Woods, Gloucestershire, England, UK, September
Rays of sunlight through mixed Beech (Fagus sp) / Oak (Quercus sp) forest after heavy downpour of hail, Peerdsbos, Brasschaat, Belgium, June 2014
Sun shining through tree at dawn. Klein Schietveld, Brasschaat, Belgium. July 2019
Sun rays filter through row of Oak (Quercus robur) trees. Peerdsbos, Brasschaat, Belgium. November
American oak (Quercus rubra) trees in avenue, sun shining through. Peerdsbos, Brasschaat, Belgium. November 2013
Sun beam at sun rise over the Grand Canyon of the Yellowstone, Yelllowstone National Park, Wyoming, USA. January 2016
Sunbeams shining through Bamboo (Bambusidae) leaves. Shunan Zhuhai National Park, Sichuan Province, China
Pyramid of sea ice catching the first rays of the returning sun in the high Arctic, Svalbard, Spitsbergen, Norway, February 2009. Taken on location for the BBC series, Frozen Planet
Dewy grass stem, backlit by rising sun, Tamar Lakes, Cornwall, UK. September 2012
RF- Moorland view from Littaford Tor, near Two Bridges, Dartmoor National Park, Devon, England, UK, October 2012
Beech (fagus) forest in autumn. Vosges mountain, France, October
Gannet (Morus bassanus) chick in a line of adults. Shetland Islands, Scotland, UK, August
RF- New Forest pony near Bratley View, New Forest, Hampshire, UK. November
Grey heron (Ardea cinerea) in rain with another heron behind, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Early morning shafts of light through deciduous forest. Bandhavgarh National Park, India
Whale shark (Rhincodon typus) front view portrait, Cenderawasih Bay, West Papua. Indonesia
Green turtle (Chelonia mydas) swimming with snorkelers in the background, Canary Islands, Spain, July
Hawksbill sea turtle (Eretmochelys imbricata) Bonaire, Leeward Antilles, Caribbean region, Netherlands Antilles
Nurse shark (Ginglymostoma cirratum) throwing up sand as it hunts in the sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Mountain forests where the Yunnan snub-nosed monkey (Rhinopithecus bieti) lives, Yunnan, China
Rainforest and valley in the Sarawaget range, Huon Peninsula, Papua New Guinea. November 2006
Silversides (Atherinomorus lacunosus) school just below surface in evening light, Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Coastal waters of the Atlantic ocean, looking toward the Treshnish islands, Scotland, UK, May
Rays of light in a fen on a misty morning, Oisterwijkse Bossen en Vennen Reserve, Oisterwijk, The Netherlands, January
Perito Moreno Glacier, Patagonia, Southern Argentina. January 2014
Stockdove (Columba oenas) shadow on the trunk of beech tree. Black Forest, Germany, April
The Quiraing, Trotternish Peninsula, Isle of Skye, Inner Hebrides, Scotland, UK. January 2015
Polar bear (Ursus maritimus) cub at entrance to den, Manitoba, Canada. March
Lagoa Misteriosa at the bottom of a sink hole in limestone region, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil November 2016. Photographed for The Freshwater Project
Brydes whale (Balaenoptera edeni) tail, Trincomalee, Eastern Province, Sri Lanka, Bay of Bengal, Indian Ocean