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Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA
Domestic rabbit, French Lop, Blue, young male, 6 months, studio portrait
Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009
Western grebe (Aechmophorus occidentalis) pair swimming, with reflections, Bolsa Chica Ecological Reserve, California, USA, February
Chilean huemul or South Andean deer (Hippocamelus bisulcus), Torres del Paine National Park, Patagonia, Chile. Endangered species
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Black bear (ursus americanus) cub standing on hind legs, Yellowstone National Park, USA, June
Leaf cutter ant (Atta sp) female worker carrying leaf to nest, with a smaller minor ant riding. The minors job is to protect her sister from parasitic flies. Costa Rica, December
Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November
Hoopoe (Upupa epops) in flight, Hungary. Multi flash, long exposure. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition
Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January
A male Spiny Seahorse (Hippocampus guttulatus) hides in seagrass. Etang De Thau (Thau Lagoon), Montpellier, France, Europe, September
West Highland White Terrier leaping
Grey Andalusian / Spanish stallion running, California, USA
Male Panther Chameleon (Furcifer pardalis) feeding on Preying mantis in beach side vegetation. Bay of Antongil, Masoala Peninsula National Park, north east Madagascar
European Mouflon (Ovis musimon) male head profile. The Netherlands, January
European Otter (Lutra lutra) walking across snow while sniffing the ground. The Netherlands, December. Captive
Domestic dog, Afghan Hound / Tazy / Baluchi Hound, head portrait
Silhouette of Brown bear (Ursus arctos) beside Lake Kuril at dawn, Kamchatka, Far east Russia, August 2008
Great white shark (Carcharodon carcharias) leaping out of the water. False Bay, South Africa, July 2010
Domestic pig, free range sow and piglet sniffing noses, UK, August 2010
Giant leaf frog (Phyllomedusa bicolor) climbing along branch in rainforest, Iwokrama Reserve, Guyana
A pair of Australian Sea Lions (Neophoca cinerea) socialising in the shallows. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January
Split level image of Caribbean reef shark (Carcharhinus perezi) breaking surface at dusk. Grand Bahama, Bahamas. Digitally manipulated image
Great Dane, black coated bitch, and Magyar Vizsla / Hungarian Pointer, tan coated maleGreat Dane, black coated bitch, and Magyar Vizsla /Hungarian Pointer, tan coated male, double portrait in profile together
A Shagya Arab stallion (Equus caballlus) is shown in hand in the courtyard of the Babolna Arabian Stud, Babolna, Komarom-Esztergom, Hungary
Green woodpecker {Picus viridis} male in flight, Lorraine, France
Trumpeter Swans (Cygnus buccinator) in winter morning mist, showing aggression during courtship behaviour, Mississippi River, Minnesota, USA, February
Galapagos green turtle (Chelonia agassizii) surfaces to breathe. Floreana Island, Galapagos Islands, Ecuador. East Pacific Ocean
Erect-crested penguins (Eudyptes sclateri) pair on coast, Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Gentoo Penguin (Pygoscelis papua) Wiencke Island, Antarctic Peninsula, Antarctica
RF - Harts tongue fern (Phyllitis scolopendrium) leaf unfurling and creating the shape of a heart, Cornwall, UK, May
African lion (Panthera leo) male shaking water off mane after rain, Masai-Mara Game Reserve, Kenya. September
Spectacled caiman (Caiman crocodilus) Rio BaiAia Bonita, Bonito, Mato Grosso do SulSpectacled caiman (Caiman crocodilus) Rio Baiia Bonita, Bonito, Mato Grosso do Sul, Brazil
Baby ground squirrel (Xerus inauris) using its tail for shade, Kgalagadi Transfrontier Park, South Africa. Non-ex
Pinocchio lizard (Anolis proboscis) male on stem, Mindo, Pichincha, Ecuador, Endangered species
Capped heron (Pilherodius pileatus) perched on branch, Yasuni National Park, Orellana, Ecuador
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
Male Bengal tiger (Panthera tigris tigris) known as Sundar (B2) patrolling territory, Bandhavgarh National Park, Madhya Pradesh, India. Endangered species
Tiger (Panthera tigris tigris), male portrait, Bandhavgarh, India, February 2013
RF- Tiger (Panthera tigris) 14-month Lakshmi cub resting in tree, Bandhavgarh National Park, India. Endangered species
Common dolphin (Delphinus delphis) surfacing, Atlantic ocean, Portugal, September
RF- Pod of Atlantic spotted dolphins (Stenella frontalis) over shallow sand bank accompanied by larger Bottlenosed dolphin (Tursiops truncatus). Sandy Ridge, Little Bahama Bank, Bahamas
Kori bustard (Ardeotis kori) male, performing mating display, silhouetted at dusk, El Karama Wilderness Conservancy, northern Kenya
Tufted duck (Aythya fuligula) on water on a misty morning, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. June
Female Cheetah (Acinonyx jubatus) standing in long grass on the savannah, Okavango Delta, Botswana, Africa
Blackbird (Turdus merula) and falling snow. Leicestershire, UK, February