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Female Bison (Bison bison) with young calf, walking over snow in front of frost-covered tree, Yellowstone National Park, USA. February
African elephant (Loxodonta africana) and Nyala (Tragelaphus angasii) at waterhole. Tembe Elephant Park. KwaZulu-Natal. South Africa
Roe deer (Capreolus capreolus) doe with fawn standing in hedgerow, Fife, Scotland, UK. September
Bison (Bison bison) being circled by wolves (Canis lupus) hunting in snow, Yellowstone National Park, USA. January
Two Red deer (Cervus elaphus) stags fighting, Bradgate Park, Leicestershire, UK. October
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd at sunrise, Masai-Mara Game Reserve, Kenya. November
Red deer (Cervus elaphus) stag jumping over a stock fence along roadside, Cairngorms National Park, Scotland, UK. February
Red deer (Cervus elaphus) stag walking through falling snow, Cairngorms National Park, Scotland, UK. January
Putorana snow sheep (Ovis nivicola borealis) lamb, standing precariously on sheer cliff face, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Three female Putorana snow sheep (Ovis nivicola borealis) climbing up rocky mountainside with lamb following, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Rainbow arching over mountains and sunlit trees in snowstorm, Northern Baikal, Siberia, Russia. October, 2020
Female Highland cow and calf (Bos taurus) next to hay bale, Exmoor National Park, Somerset / Devon, England. November
Female Highland cow (Bos taurus) licking its calf next to hay bale, Exmoor National Park, Somerset / Devon, England. November
Red deer stag (Cervus elaphus) with female during rut, Exmoor National Park, Somerset / Devon, England. November
Two Cheetah (Acinonyx jubatus) hunting Blue wildebeest (Connochaetes taurinus). Masai Mara national reserve, Kenya, East Africa
White-tailed deer (Odocoileus virginianus) doe with fawn in woodland, Acadia National Park, Maine, USA. July
Young White-tailed deer (Odocoileus virginianus), alert, looking back towards photographer, Finland. September
Male Moose (Alces alces) with antlers, portrait, Finland. September
Mother Moose (Alces alces), with calf, grazing in meadow, Finland. September
Two Reindeer (Rangifer tarandus tarandus) in snowy landscape, Finland. April
Mother White-tailed deer (Odocoileus virginianus) allowing young to suckle, Finland. September
Two male Alpine Ibex (Capra ibex) fighting on a mountain slope at dawn, Varaita Valley, Alps, Italy, July
Red deer (Cervus elaphus) stag amongst ferns, Bradgate Park, Leicestershire, UK. October
Red deer (Cervus elaphus) stag bellowing amongst bracken, Bushy Park, London, England, UK, October
Marsh deer (Blastocerus dichotomus) walking through vegetation, Pocone, Brazil
Bull Moose (Alces alces) drinking from mountain stream, Grand Teton National Park, Wyoming, USA. October
Male Saiga antelope (Saiga tatarica) rutting in winter, The Black Lands National Park, Kalmykia, Russia
Female Cuviers gazelle (Gazella cuvieri) calf, age 6 months, with its mother, Djebel Ouarkziz, Southern Morocco, Africa. Customized camera-trap
Moose (Alces alces) two bulls sparring in willow flat grasslands, Grand Teton National Park, Wyoming, USA. October
Fallow deer (Dama Dama) stags in velvet standing in a shady woodland spot, Knepp Estate, Sussex, UK, June
Herd of domestic Water buffalo (Bubalus arnee) in dust at dusk, Lake Kerkini area, Greece, July
Alpine ibex (Capra ibex) adult males fighting on a steep mountain side, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Roe deer (Capreolus capreolus) feeding in field with flowering Poppies (Papaver rhoeas) and Cornflower (Centaurea cyanus), De Inslag, Brasschaat, Belgium, June
Himalayan ibex (Capra sibirica hemalayanus) males, resting and grazing. They live at elevations of 3800m and higher, western Himalaya mountains, Kibber Wildlife Sanctuary, India. April
Red deer (Cervus elaphus) stags sparring in the frost. Richmond Park, London, England, UK. December
American buffalo (Bison bison) herd. Yellowstone National Park, Wyoming, USA
Red deer (Cervus elaphus) stag standing in bracken. London, UK. September
European mouflon (Ovis musimon) male, Sierra de Andujar, Andalusia, Spain. January
Southern chamois (Rupicapra pyrenaica) male, Picos de Europa, Spain, August
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Northern Serenget National Park, Tanzania (early September)
Angolan giraffes (Giraffa giraffa angolensis) approaching a scarce waterhole. Etosha National Park, Etosha, Namibia
European Bison (Bison bonasus) in winter, Białowieza National Park, Poland. January
Red deer (Cervus elaphus) in the pine forest bellowing during the rutting season. Barycz Valley Landscape Park, Poland. September
White-tailed deer (Odocoileus virginianus), two fawns playing in field. Kirkkonummi, Finland. September
Red deer (Cervus elaphus) stags fighting during the rutting season, Bushy Park, London, UK October
White-lipped deer (Cervus albirostris) walking on the Tibetan plateau, Serxu, Garze Prefecture, Sichuan Province, China
RF - Red deer (Cervus elaphus) stag standing amongst Scots pine (Pinus sylvestris) tree trunks in forest. Cairngorms National Park, Scotland, UK. January
Blue sheep (Ovis nahoor) Angsai Nature Reserve (Valley of the Cats), Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China