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Iberian ibex (Capra pyrenaica), adult male, in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. February
Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September
Red deer (Cervus elaphus) stag bellowing, Richmond Park, London, England, UK. October
Saiga antelope (Saiga tatarica) male, standing in grassland in morning sunlight, Betpak Dala, Kazakhstan. Critically endangered
Domestic yak (Bos grunniens) lying under prayer flags near shores of Nam-Tso Lake (at 4, 730m), second largest salt lake in China. Tibet, China
Chamois (Rupicapra rupicapra) head portrait, Schwyz, Switzerland. November
Ibex (Capra ibex) feeding on grass, head portrait, Schwyz, Switzerland. August
Two Ibex (Capra ibex) males, standing on hillside, Lucerne, Switzerland. August
Iberian ibex (Capra pyrenaica), pair with male top right, during rutting season. Sierra Nevada National Park, Andalusia, Spain. October
Iberian ibex (Capra pyrenaica) kid peering over rocks. Sierra Nevada National Park, Andalusia, Spain. June
Iberian ibex (Capra pyrenaica) yearling making its way down snow covered slope. Sierra Nevada National Park, Andalusia, Spain. February
Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), female, being pestered by kid, aged two months, begging for milk. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica) kid, aged two months, making its way down cliff, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), female, grazing, being pestered by kid, aged two months, begging for milk, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Four Iberian ibex (Capra pyrenaica) kids, aged two months, playing, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), adult male, sitting down in snow covered landscape during snowfall. Sierra Nevada National Park, Andalusia, Spain. April
Iberian ibex (Capra pyrenaica), alert female with kid aged between one and two months. Sierra Nevada National Park, Andalusia, Spain. May
Iberian ibex (Capra pyrenaica), adult female, walking along coastal cliff. Mediterranean coast, Andalusia, Spain. November
South Andean deer (Hippocamelus bisulcus), adult female with fawn, resting on mountainside. Cerro Castillo National Park, Patagonia, Chile. April
Red deer (Cervus elaphus), young male, swimming through water. Bushy Park, London, UK. September
Masai giraffe (Giraffa camelopardalis tippelskirchi) running across plain. Amboseli National Park, Kenya. July. Digitally enhanced
Nilgiri tahr (Nilgiritragus hylocrius) standing next to road signage meant to warn vehicles about the possible road crossings of tahrs in the area, Valparai, Tamil Nadu, India
Springbok (Antidorcas marsupialis) herd in grassland after rain, Palmwag Conservancy, Damaraland, Namibia
Gemsbok (Oryx gazella) herd in desert after the wet season with Naukluft Mountains in the background, Namib-Naukluft National Park, Namibia
Gemsbok (Oryx gazella) pair standing in grass after wet season in desert, Namib-Naukluft National Park, Namibia
Gemsbok (Oryx gazella) female and calf walking past sand dune, Sossusvlei UNESCO World Heritage Site, Namib desert, Namib-Naukluft National Park, Namibia
Sika deer (Cervus nippon) mother walking along hilltop as fawn climbs slope between rocks and trees, with mountain forest behind, Land of the Leopard National Park, Russian Far East
Sika deer (Cervus nippon) doe standing on rocky outcrop overlooking mountain forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. January
Sika deer (Cervus nippon) stag wading through river in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) herd feeding beside pool in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) doe with fawn standing on rocky outcrop overlooking forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) stag walking on mountain slope at the edge of woodland, with forested mountains in the background, Land of the Leopard National Park, Russian Far East
Sika deer (Cervus nippon) herd walking past fallen tree in forest as one looks around, Land of the Leopard National Park, Russian Far East. Taken with remote camera. June
Sika deer (Cervus nippon) doe walking across snow covered mountain slope, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
White-tailed deer (Odocoileus virginianus) fawn, standing among wildflowers and long grass, Acadia National Park, Maine, USA. June
White-tailed deer (Odocoileus virginianus) reaching up to feed on leaves, Acadia National Park, Maine, USA. June
Alpine ibex (Capra ibex) male, with kid on mountain ridge in heavy winter snowfall, Valsavarenche, Gran Paradiso National Park, Italy. January
Bull moose (Alces alces) crossing a mountain creek at sunset, Grand Teton National Park, Wyoming, USA. October
Spotted deer (Axis axis), small herd leaping through grass, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis) herd in the forest, with just their antlers visible. Bardia National Park, Terai, Nepal
Two Red deer (Cervus elaphus) stags, fighting during the rut, Cheshire, UK. July
Frog (Anura) in waterhole at night with two Cape buffalo (Syncerus caffer) in background, Zimanga Game Reserve, KwaZulu-Natal, South Africa
Aerial view of migrating Wildebeest (Connochaetes taurinus) on the border of Ngorongoro Conservation Area and Serengeti National Park, Tanzania, Africa
Red Lechwe (Kobus leche) herd crossing swamp surrounding a small island, Okavango Delta, Botswana, Africa
Red Lechwe (Kobus leche) herd resting on a small island in swamp, Okavango Delta, Botswana, Africa
Two Red deer (Cervus elaphus) standing alert under a tree in woodland. The last surviving indigenous herd of Red deer in Republic of Ireland has been present in Killarney Valley since neolithic
Giraffe (Giraffa camelopardalis angolensis) calf, aged 6 weeks, with two Yellow-billed oxpeckers (Buphagus africanus) perched on its head feeding on parasites, Okavango Delta, Botswana, Africa
