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Gemsbok (Oryx gazella) in sand dunes, Namibia. Finalist in the Mammals Category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
American bison (Bison bison) female covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
RF - Bison (Bison bison) breathing in the cold air, Yellowstone National Park, USA, February
Reindeer (Rangifer tarandus) herd, antlers in velvet, walking across upland moor, Cairngorms, Scotland, UK, August
Herd of American bison (Bison bison) in snow, Yellowstone National Park, Wyoming, Yellowstone, January
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Red deer (Cervus elaphus) stag in upland landscape. Lochcarron, Highlands, Scotland, UK
Okapi (Okapia johnstoni). Beauval Zoo Parc, France. Captive
Male Saiga antelope (Saiga tatarica) in winter, The Black Lands National Park, Kalmykia, Russia
Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October
North American Bison (Bison bison) coated in frost standing on snow, Yellowstone National Park, Wyoming, USA, January
Caribou / Reindeer bull (Rangifer tarandus) close-up grazing, Denali National Park, Alaska, USA, September
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
American bison (Bison bison) Yellowstone National Park, Montana, USA
Alpine ibex (Capra ibex) female, Jura, Switzerland, September
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Siberian musk deer (Moschus moschiferus) male in snow, Irkutsk, Russia. January
Impala with Oxpeckers. Kruger National Park, South Africa
American bison (Bison bison) male walking through deep snow. Hayden Valley. Yellowstone National Park, Wyoming, USA. January
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
Gaur (Bos gaurus) Manas National Park UNESCO World Heritage Site, Assam, India
Aerial view of Reindeer (Rangifer tarandus) fording water whilst being herded by Nenet herders. Nenets Autonomous Okrug, Arctic, Russia, July 2017
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Roe Deer stag (Capreolus capreolus) standing in a woodland glade. Potterick Carr Nature Reserve, Doncaster, Yorkshire, England, UK, July
Two Red deer (Cervus elaphus) stags, fighting during the rut, Cheshire, UK. July
Roe deer (Capreolus capreolus) doe, head portrait, Fife, Scotland, UK. July. Captive
Red deer stag (Cervus elaphus) during rut on lookout, Exmoor National Park, Somerset / Devon, England. October
Female American buffalo (Bison bison) walking across grassland with newborn calf, Yellowstone National Park, Wyoming, USA. May
Male Nubian Ibex (Capra nubiana) at the Arabian Wildlife Centre captive-breeding project, Sharjah, United Arab Emirates. (captive)
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Alpine ibex (Capra ibex) resting in mountain pasture, Ticino, Switzerland
Tibetan gazelle / Goa (Procapra picticaudata) Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
Autumn quaking aspen grove (Populus tremuloides) with trunks scarred by browsing elk (Cervus canadensis). Below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
White-tailed deer (Odocoileus virginianus) doe standing in snow, New York, USA
Altai argali sheep (Ovis ammon), three grazing on slope. Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China. October
Tibetan gazelle (Procapra picticaudata) in snow. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October
Chital (Axis axis) female leaping. Tadoba Andhari Tiger Reserve / Tadoba National Park, Maharashtra, India
Red deer (Cervus elaphus) silhoutted at dusk, Richmond Park, London, England, UK. October
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Moose (Alces alces) bull swimming across river. Grand Teton National Park, Wyoming, USA. September
Silver Dik-dik (Madoqua piacentinii) portrait, Masai Mara, Kenya. March
Gayal / Mithun (Bos frontalis) on river bank. Arunachal Pradesh, North East India. November 2014
RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Chinese Water Deer (Hydropotes inermis) Norfolk UK September
Roe deer (Capreolus capreolus) buck with vegetation on antlers in rutting season, Scotland, UK. August
Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Alpine ibex (Capra ibex) struggling in deep snow on a steep slope, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, January
Snow leopard (Uncia uncia) with Bharal (Pseudois nayaur) prey, Serxu County, Garze Prefecture, Sichuan Province, China
Pere Davids deer / Milu (Elaphurus davidianus) during the rutting season, stag, Hubei Tian ezhou Milu National Nature Reserve, Shishou, Hubei, China
Red deer (Cervus elaphus) stag, Saxony, Germany. September
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Sable antelope (Hippotragus niger) bull charging through the shallows of the Selinda Spillway, with another behind, northern Botswana, August
Group of four Giraffes (Giraffa camelopardalis) Mkuze, South Africa
Moose (Alces alces) female with twin calves, Baxter State Park, Maine, USA, June
Asiatic Wild Water Buffalo (Bubalus arnee), Royal Manas National Park, Bhutan
Red deer (Cervus elaphus) male stag roaring portrait, rut season, Jura, Scotland, UK, September
Masai giraffe (Giraffa camelopardalis tippelskirchi) running across plain. Amboseli National Park, Kenya. July. Digitally enhanced
Putorana snow sheep (Ovis nivicola borealis) lamb, standing precariously on sheer cliff face, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Male Moose (Alces alces) with antlers, portrait, Finland. September
Snow sheep (Ovis nivicola borealis) adult with lamb, huddled against the rockface, Plateau Putorana Nature Reserve, Siberia, Russia
Two male Alpine Ibex (Capra ibex) fighting on a mountain slope at dawn, Varaita Valley, Alps, Italy, July
Rothschild giraffes (Giraffa camelopardalis rothschildi), three walking in shade and three walking in light, Mara North Conservancy, Kenya. October
Bull Moose (Alces alces) drinking from mountain stream, Grand Teton National Park, Wyoming, USA. October
Red deer stag (Cervus elaphus) rear view, shaking water off itself after the rain, Bradgate Park, Leicestershire, UK
Moose (Alces alces) bull crossing mountain river at sunset, Grand Teton National Park, Wyoming, USA
Herd of Salers cattle - cows and calves (Bos sp. ), Aisne, France, MayHerd of Salers cattle - cows and calves (Bos sp.), Aisne, France, May
Alpine ibex (Capra ibex) adult males fighting on a steep mountain side, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Roe deer (Capreolus capreolus) feeding in field with flowering Poppies (Papaver rhoeas) and Cornflower (Centaurea cyanus), De Inslag, Brasschaat, Belgium, June
Bharal / Blue sheep (Pseudois nayaur) herd standing on a rocky slope, higher Himalaya mountains, Kibber Wildlife Sanctuary, India. March
Fallow deer (Dama dama) hind and fawn in early morning mist. London, UK. September
Southern chamois (Rupicapra pyrenaica) male, Picos de Europa, Spain, August
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Northern Serenget National Park, Tanzania (early September)
South African / Cape giraffe (Giraffa camelopardalis giraffa), two males necking / mock fighting, looking like a double headed animal, Zimanga Private Game Reserve, Zululand, South Africa
Red deer stag (Cervus elaphus), Bushy Park, London, UK October
Red deer (Cervus elaphus) stags fighting during the rutting season, Bushy Park, London, UK October
Fallow deer (Dama dama) white stag on frosty morning, Bushy Park, London, UK. November
Taiwan / Formosan Reevess muntjac baby, (Muntiacus reevesi formosanus), Walami cabin, Yushan National Park, Taiwan. Endemic subspecies
Tibetan antelope / Chiru (Pantholops hodgsonii) three silhouetted, Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
