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Lichen (Xanthoria parietina) found growing on a twig, showing mass of apothecia (cup-shaped fruiting bodies). Peak District National Park, UK. January
Rowan (Sorbus aucuparia) fresh leaves in spring, Beinn Eighe NNR, Highlands, NW Scotland, UK, May
Panther Chameleon {Furcifer pardalis} close-up of skin, green colouration. Masoala Peninsula National Park, north east Madagascar
Panther chameleon {Furcifer pardalis} walking along branch, Masoala Peninsula National Park, north east Madagascar
Fly agaric fungus (Amanita muscaria) cap detail. Peak DIstrict National Park, Derbyshire, UK. October
RF- Alpine marmot (Marmota marmota) reaching upwards, Hohe Tauern National Park, Austria. July
Male African lion in his prime, portrait {Panthera leo} Chobe NP, Botswana
Grey heron (Ardea cinerea) beneath waterfall. Ambleside, Lake District, UK, November. Highly commended, Habitat category, British Wildlife Photography Awards (BWPA) competition 2012
Grizzly bear (Ursus arctos horribilis) in grassland, Yellowstone National Park, Wyoming, USA, October
Cape buffalo (Syncerus caffer) crossing the Chobe River, followed by swarm of flies, Chobe Game Reserve, Botswana, Africa
Meerkat (Suricata suricatta) group standing apart, Kgalagadi National Park, South Africa
European Black vulture (Aegypius monachus) portrait of two birds, Campanarios de Azaba Biological Reserve, a rewilding Europe area, Salamanca, Castilla y Leon, Spain
White-backed Woodpecker (Dendrocopos leucotos). Bieszczady, Carpathian Mountains, Poland, February
Portrait of a wild Carpathian Grey Wolf (Canis lupus lupus) in snow-bound woodland habitat. Bieszczady, Carpathian Mountains, Poland, December
Southern white rhinoceros (Ceratotherium simum simum) grazing. Ol Pejeta Conservancy, Kenya, 2009. Threatened species
African elephant (Loxodonta africana) herd, Tarangire NP, Tanzania
View of lake surrounded by sand dunes, with some vegetation growing around the edges, Namib Naukluft NP, Namib desert, Namibia
Hippopotamus and calf {Hippopotamus amphibius} Virunga NP, Congo
Black-bellied Plover (Pluvialis squatarola) chicks in the nest, seen from overhead, Yukon Delta National Wildlife Refuge, Alaska, USA June
Fer-de-lance (Bothrops asper) camouflaged on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Satanic leaf-tailed gecko (Uroplatus phantasticus) on twig, Andasibe-Mantadia National Park, Alaotra-Mangoro Region, Madagascar
Merveille-du-Jour moth (Dichonia aprilina), camouflaged on lichen. The National Forest, Leicestershire, UK. September
Parsons chameleon {Calumma parsonii} female walking along vine in tropical rainforest habitat. Andasibe-Mantadia NP, Madagascar
Bornean horned frog (Megophrys nasuta) amongst leaf-litter on forest floor. Danum Valley, Sabah, Borneo, Malaysia
Melanistic leopard / Black panther (Panthera pardus) on territorial patrol on track, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Silky shark (Carcharhinus falciformis), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January. Vulnerable
Waterfall at mouth of Kitum cave Mt. Elgon National park, Kenya, August 2017
Spot-bellied eagle owl (Bubo nipalensis) perched on branch, Jim Corbett National Park, Uttarakhand, India
American crocodile (Crocodylus acutus), underwater, Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January. Vulnerable species
Cheetah (Acinonyx jubatus) cubs 4 months, Masai-Mara Game Reserve, Kenya. Vulnerable species
Cheetah (Acinonyx jubatus) mother carrying its cub aged 4 to 5 weeks, Masai-Mara Game Reserve, Kenya. Vulnerable species
Wildebeest (Connchaetes taurinus) running down bank of the Mara River during migration, Masai Mara, Kenya
Leopard (Panthera pardus) female yawning, Samburu game reserve, Kenya
Young Meerkat (Suricata suricatta) standing on its hind legs near the edge of Makgadikgadi Pans National Park, Botswana, April
Male Caucasian turs (Capra caucasica) rutting with Dykh-tau Mountain in the background, Kabardino-Balkarsky Zapovednik / Reserve, Russia, July
Two curious young Polar bears (Ursus maritimus), Barter Island, off the 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska, October 2011
Portrait of Polar bear (Ursus maritimus) cub along the Bernard Spit during autumn freeze up, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska, October 2011
Silverback Mountain gorilla (Gorilla beringei) in the bamboo forest, this is Munyinya the leader of Hirwa group, Sabyinyo volcano, Volcano National Park, Rwanda
Tres Marias Rabbit (Sylvilagus graysoni). Maria Madre Island, Islas Marias Biosphere Reserve, Sea of Cortez (Gulf of California), Mexico, July. Endangered
Mountain pansy (Viola lutea) flowers, Strathspey, Cairngorms National Park, Scotland, June
Dall sheep (Ovis dalli) yearling lamb running across alpine tundra, Denali National Park, Alaska, USA
Blue ice melt pool in snow near the peak of La Munia Circo. The Pyrenees, Aragon, Spain, May 2011
Four Wisent / Bison (Bison bonasus) standing in a row. Bialowieza Forest, Bialowieza National Park, Poland, November
Woody / Monkey ladder vine (Bauhinia glabra) Palo Verde National Park, Costa Rica
Palm trees (Licuala valida) in swamp forest, Lambir NP, Borneo, Sarawak, Malaysia
Silver Birch (Betula verrucosa) reflected in Loch Beinn a Mheadhoin. Glen Affric National Nature Reserve, Scotland, December
Black bear (Ursus americanus) cub, brown phase, looking out from behind tree trunk, Yellowstone National Park, Wyoming, USA, May
White sand dunes against blue sky, White Sands National Monument, New Mexico, USA, February 2009