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RF - Common blue butterflies (Polyommatus icarus) resting on grass. Vealand Farm, Devon, UK. September
Pectoral sandpiper (Calidris melanotos) juvenile feeding around edge of shallow flood, Llanrhystud, Ceredigion, West Wales, UK, September
Water vole (Arvicola amphibius) Kent, UK September
Spiny / Yellow Seahorse (Hippocampus guttulatus). Studland Bay, Dorset, UK, September
Pine Marten (Martes martes) standing alert on a trunk. Cairngorms National Park, Scotland, September
Red Squirrel (Sciurus vulgaris), at woodland pool, Cairngorms National Park, Scotland, UK.November
White rhinoceros (Ceratotherium simum) male and female, face to face, Mosi Oa Tunya National Park, Zambia
White rhinoceros (Ceratotherium simum) with a long horn, Marakele National Park, South Africa
Brown hare (Lepus europaeus) adult in arable field, Scotland, August
Red squirrel (Sciurus vulgaris) standing on log surrounded by heather, Cairngorms National Park, Scotland, UK, August
Meadow Brown Butterfly (Maniola jurtina) feeding from Scabious flower (Scabiosa) North Downs, Surrey, England, UK, August
Clouded yellow butterfly (Colias croceus) on Fleabane. Dorset, UK, August
Brimstone butterfly (Gonepteryx rhamni) on Devil s-bit Scabious (Succisa pratensis). Peak District National Park, Derbyshire, UK. August
Welsh Ponies (Equus caballus) engaged in mutual grooming, Llanbedr Hill, Powys, Wales, August
Lesser Marsh Grasshopper (Chorthippus albomarginatus) brown and green form Lewisham, London, August
Grey heron (Ardea cinerea) head profile, Regents Park, London, UK August
Red-necked phalarope (Phalaropus lobatus) female about to strike at a fly hovering nearby. Shetland Islands, Scotland, UK, August (non-ex)
Golden plover (Pluvialis apricaria) profile portrait on open moorland in evening light. Shetland Islands, Scotland, UK, August (non-ex)
Dunlin (Calidris alpina) portrait. Vardo, Finnmark, Norway. May
Exmoor pony {Equus caballus} grazing amongst flowering heather {Ericaceae}, near Porlock Hill, Exmoor National Park, Somerset, UK. August
Pileated Woodpecker (Dryocopus pileatus) male, at feeding excavation in tree trunk in winter, Freeville, New York, USA, January
American oystercatchers (Haematopus palliatus) courting pair performing Piping Display, calling as they walk along together, Fort DeSoto Park, Florida, USA, March
Puffin (Fratercula artica) with orange sea lichen (Caloplaca marina) covered roch behind, Shiant Isles, Outer Hebrides, Scotland, UK. June
Red squirrel (Sciurus vulgaris) feeding in snow. Scotland, UK. March
Rivolis hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica
Hedgehog (Erinaceus europaeus) walking through puddle. Sado Estuary, Portugal. March
Flightless cormorant (Phalacrocorax harrisi), adult and two chicks. Beagle Crater, Isabela Island, Galapagos
Southern lechwe (Kobus leche), two males about to fight, facing each other. Okavango Delta, Botswana
Thomsons gazelle (Eudorcas thomsonii) male and two females standing in rainstorm, Masai Mara Game Reserve, Kenya
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Adult Kea (Nestor notabilis) close-up portrait. Arthurs Pass National Park, New Zealand, July. Endangered Species
Madagascar / Malagasy kingfisher (Alcedo vintsioides) perching, Anjajavy Private Reserve, north west Madagascar
Komodo dragon (Varanus komodoensis) Rinca Island, Indonesia
Chamois (Rupicapra rupicapra) in the snow, Jura Switzerland, November
Thorny dragon (Moloch horridus) in desert habitat, Australia
Head portrait of black Andalusian stallion and mare meeting for the first time in southern Spain, Europe
Black korhaan (Eupodotis afra) male displaying and calling. Etosha National Park, Namibia, March
African buffalo (Syncerus caffer) feeding on water lillies in Chobe River, Chobe National Park, Botswana. May
Honey bee (Apis mellifera) on comb with honey, Kiel, Germany, June
Llama standing in mud at the edge of Laguna Colorada, Bolivia
Blue cheeked bee eater (Merops persicus) two squabbling, Oman, April
Osprey (Pandion haliaetus) male and female head to head at nest, just before mating, Finland April
Nile crocodile (Crocodylus niloticus). Zimanga Private Game Reserve, KwaZulu-Natal, South Africa. September
Bignose unicornfish (Naso vlamingii) profile of male displaying (the blue stripe at the bottom of the frame is the tail of another individual) Baa Atoll, Maldives. Indian Ocean
Black stork (Ciconia nigra) Hungary May
Spiny predatory katydid (Listroscelis sp) photographed on a white background in mobile field studio, Osa Peninsula, Costa Rica
RF - Eastern tiger swallowtail butterfly (Papilio glaucus) nectaring on Zinnia in farm garden, wild and free. Essex, Connecticut, USA
Svalbard reindeer (Rangifer tarandus platyrhynchus) with bloody antlers shedding velvet, Spitsbergen, Svalbard, Norway, September
