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Feral domestic cattle, Puerto Fragata, Isabela Island, Galapagos
Buff-tailed bumblebee (Bombus terrestris) queen about to land at her nest burrow in a garden lawn with loaded pollen sacs to provision grubs that will become future workers for her colony, Wiltshire
Oriental Magpie Robin (Copsychus saularis) female, Whitefield, Bangalore, India, March
American crocodile (Crocodylus acutus), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
Great anglehead lizard (Gonocephalus grandis) male, Tioman Island, Malaysia
Hippopotamus (Hippopotamus Amphibius) with back covered in water hyacinth on back after leaving water. Mana Pools National Park Zimbabwe
Tibetan gazelle or Goa, (Procapra picticaudata) leaping, in mid air. Dulan Nature Reserve, Tibetan Plateau, Qinghai, China. October
Chital (Axis axis) female leaping. Tadoba Andhari Tiger Reserve / Tadoba National Park, Maharashtra, India
Pine grosbeak (Pinicola enucleator), perched by berries in Rowan tree, Finland, January
Wood warbler (Phylloscopus sibilatrix) amongst willow catkins, Finland, May
Common loon (Gavia immer), on a lake, Michigan, USA. June
Garibaldi (Hypsypops rubicundus) in forest of giant kelp, (Macrocystis pyrifera) off Santa Barbara Island, California, USA
Bullfinch (Pyrrhula pyrrhula) in flight, Kuusamo, Finland, March
Moose (Alces alces) bull, portrait. Grand Teton National Park, Wyoming, USA. September
Carters semaphore gecko (Pristurus carteri) standing on rocky ground. Oman, June
Puma (Puma concolor puma), young male crossing road in front of car. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December 2018
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Verreauxs sifaka lemur (Propithecus verreauxi) dancing or skipping across open ground, Gallery Forest, Berenty Reserve, southern Madagascar
African penguin (Spheniscus demersus) standing on rock at dusk, with clustered penguins in background. South Africa. November
Greater flamingo (Phoenicopterus roseus) pair at rest in water, Cape Town, South Africa, July
Grey peacock-pheasant (Polyplectron bicalcaratum) Tongbiguan Nature Reserve, Dehong, Yunnan, China
Temmincks tragopan pheasant (Tragopan temminckii) Chongqing region, Sichuan, China
Kalij pheasant (Lophura leucomelanos) Tongbiguan Nature Reserve, Dehong, Yunnan, China
RF - Guanaco (Lama guanicoe) standing in front of mountain towers of Paine. Torres del Paine National Park, Patagonia, Chile. December 2018
False gharial (Tomistoma schlegelii) resting on rock in Mpassa river. Bateke Plateau National Park, Gabon
Springtail (Dicyrtomina ornata) on oak tree leaf, Niedersechsische Elbtalaue Biosphere Reserve, Elbe Valley, Lower Saxony, Germany, December
Woodland Kingfisher (Halcyon senegalensis) perched in tree, Serengeti, Tanzania
Trumpeter hornbill (Bycanistes bucinator) feeding, captive
Collared peccary (Pecari tajacu) standing on the rainforest floor. Madre de Dios, Peru. March
Golden-collared Tanager (Iridosornis jelskii), in the Peruvian cloud forest. Wayqecha Biological Research Station, Peru. April. Cropped
Bull Skink (Liopholis multiscutata) in coastal dunes at Browns Beach on Kangaroo Island, South Australia, Autumn
RF - Basking shark (Ceterhinus maximus) feeding on plankton in Senen Cove, Cornwall, UK. June. (This image may be licensed either as rights managed or royalty free.)
Indian peafowl or blue peafowl (Pavo cristatus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Chamois (Rupicapra rupicapra) Ballons des Vosges Regional Natural Park, Vosges Mountains, France, October
Alpine ibex (Capra ibex) on a mountain side in autumn with European larch tree (Larix decidua) trees in the back the Alps, Gran Paradiso National Park, Italy. November
Eurasian red squirrel (Sciurus vulgaris) reflected in water, Cairngorms National Park, Scotland, United Kingdom, October
Lapwing (Vanellus vanellus) Titchwell RSPB Reserve, Norfolk, England, UK. November
Grey squirrel {Sciurus carolinensis} adult feeding among the fallen leaves of Horse Chestnut tree, Derbyshire, UK
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Marine iguana (Amblyrhynchus cristatus) on coast, Galapagos
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Galapagos hawk (Buteo galapagoensis) landing, Galapagos, Ecuador. Vulnerable species
Yellow breasted antipitta (Grallaria flavotincta) Angel Paz reserve, Mindo region, Andean cloud forest, Ecuador
Plate billed mountain toucan (Andigena laminirostris) Bellavista cloud forest private reserve, 1700m altitude, Tandayapa Valley, Andean cloud forest, Ecuador
James flamingo / Puna flamingo (Phoenicoparrus jamesi). Lago Colorado, Bolivia. March
Yellow warbler (Setophaga petechia aureola) on lava rocks. Isabela Island, Galapagos. May
Mining bee (Andrena sp. ), harvesting pollen from anthers of Cherry tree (Prunus spMining bee (Andrena sp.), harvesting pollen from anthers of Cherry tree (Prunus sp.) Ripon Wisconsin USA, May
Atlantic bluefin tuna (Thunnus thynnus) off Antigonish, Nova Scotia, Canada. October