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Grey wolf (Canis lupus) standing among trees in snow-covered forest, Finland. November
American pine marten (Martes americana) standing on hind legs in deep snow, Yellowstone National Park, USA. January
Robin (Erithacus rubecula) perched on fallen apple in snow, Bishopswood, Somerset, UK. January
Moorhen (Gallinula chloropus) on water in falling snow, near Bourne, Lincolnshire, England, UK. March
Eurasian blue tit (Cyanistes caeruleus) perched on snow-covered grass, puffed up to keep warm, Białowieza National Park, Poland. January
Two European goldfinches (Carduelis carduelis) fighting over food in snow, Wrocław, Lower Silesia, Poland. February
Blackbird (Turdus merula) male eating winfall apples in snow, Broxwater, Cornwall, UK. December
Mourning dove (Zenaida macroura) perched on branch in snow storm; Milford, Connecticut, USA. February
Collie standing in snowy, frozen marsh, Eagle River, Alaska, USA. April
Sand cat (Felis margarita) portrait. Captive, occurs in North Africa, the Arabian Peninsula, Pakistan and the Middle East
Humpback whale (Megaptera novaeangliae) swimming, Tubuai, French Polynesia, Pacific Ocean
Tiger shark (Galeocerdo cuvier) swimming over sandy seabed, Tiger Beach. Bahamas, Atlantic Ocean
Mount Lewis spiny crayfish (Euastacus fleckeri) in defensive posture with claw raised, at edge of mountain stream, Mount. Lewis, Wet Tropics World Heritage area, Queensland, Australia. Endangered
Caterpillar, probably Cholla moth caterpillar, (Euscirrhopterus cosyra) on cactus, Texas, USA. May
Dusky shark (Carcharhinus obscurus) swimming through light rays during sardine run. Wild Coast, Eastern Cape, South Africa. June
Glasswing butterfly (Greta oto) resting on leaf. Captive
Long-tailed potoo (Nyctibius aethereus) camouflaged on roosting perch in lowland rainforest, Manu Biosphere Reserve, Peru
Coyote (Canis latrans) pouncing, hunting technique in Yellowstone National Park, Wyoming, USA
Cairns birdwing butterfly (Ornithoptera euphorion) male resting on fern, large butterfly endemic to wet tropics in far north Queensland, Cairns, Queensland, Australia
Mandarin duck (Aix galericulata) male swimming with autumn colours reflected in the water, London, UK, November
Thorny devil stick insect (Eurycantha calcarata), Willaumez Peninsula, New Britain, Papua New Guinea, December
Grey triggerfish (Balistes capriscus) portrait, Tenerife, Canary Islands. August
Two male Indian Wild Asses (Equus hemionus khur), looking like they share a single head, Little Rann of Kutch, Gujarat, SW India
Fallow deer (Dama dama) white stag on frosty morning, Bushy Park, London, UK. November
Taiwan / Formosan yellow-throated marten (Martes flavigula chrysospila) Alishan National Recreational Forest, Taiwan
Close up at the dental plates of Three color parrotfish (Scarus tricolor) as it comes in for a bite of hard coral, Fiji
Taiwan yellow tit ( Machlolophus holsti ) Lalashan National Forest Reserve, Taiwan. Endemic
Guanaco (Lama guanicoe) in grassland with Torres del Paine rock towers in background. Torres del Paine National Park, Patagonia, Chile. January 2020
Peregrine falcon (Falco peregrinus) in flight with open beak. The Netherlands. May
Coarse chameleon (Trioceros rudis) profile, Volcanoes National Park, Rwanda
A portrait of a clown triggerfish (Balistoides conspicillum) on a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Misool Marine Protected Area. Ceram Sea. Tropical West Pacific Ocean
A male pajama cardinalfish (Sphaeramia nematoptera) incubating eggs in his mouth (not clearly visible). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Apple hawk-moth (Langia zenzeroides formosana) Nzaus Mamei Jianshih Village, Hsssinchu County, Taiwan, Controlled conditions
Stellers sea eagle, (Haliaeetus pelagicus), on sea pack ice, Okhotsk Sea, Rausu, Hokkaido, Japan
Palm Cockatoo (Probosciger aterrimus) feeding on Beach Almond (Terminalia catappa) Lockhart River Foreshore, Lockhart River, Cape York Peninsula, Australia
RF- Northern Cardinal (Cardinalis cardinalis) male perched on branch of flowering Eastern Redbud (Cercis canadensis). Dinero, Lake Corpus Christi, South Texas, USA
TemminckAśs tragopan (Tragopan temminckii) male during courtship displayTemminck's tragopan (Tragopan temminckii) male during courtship display, showing red-blue lappets. Tibet, China
Arabian gazelle (Gazella gazella), Jaluni, Oman, May
Whooper swan (Cygnus cygnus) in snow. Jyvaskyla, Central Finland. March
Bullfinch (Pyrrhula pyrrhula) in flight, Kuusamo, Finland, March
Siberian tiger (Panthera tigris altaica) at night, taken with remote camera in Land of the Leopard National Park, Far East Russia
Brown skua (Stercorarius antarcticus), Ronge Island, Antarctica
Pin-tailed whydah (Vidua macroura) male in display flight. Allahein River, Gambia
Fire-tailed sunbird (Aethopyga ignicauda) perched amongst blossom. North Sikkim, India. April
Black rhino (Diceros bicornis) with very long horn, Lewa Wildlife Conservancy, Laikipia, Kenya. October
Brimstone butterfly (Gonepteryx rhamni) resting among foliage, Meeth Quarry, Devon, UK. June
Shoebill stork (Balaeniceps rex) eating a fish in the swamps of Mabamba, Lake Victoria, Uganda
Bactrian camel (Camelus bactrianus) male silhouetted at sunset, living in the wild but owned by a camel herdsman, Turpan Basin, Gobi Desert, Xinjiang, China