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Pixy Hawkfish (Cirrhitichthys oxycephalus) Lembeh Strait, North Sulawesi, Indonesia
Banggai Cardinalfish (Pterapogon kauderni), a shoal sheltering against Anchor Coral (Euphyllia ancora). Lembeh Strait, North Sulawesi, Indonesia
Cuckoo wrasse (Labrus mixtus) male in front of Red sea fan, (Paramuricea clavata), Stupiste In dive site, Vis Island, Croatia, Adriatic Sea, Mediterranean
Common anemonefish (Amphiprion perideraion) associated with the anemone (Heteractis magnifica) Yap, Micronesia
Sheephead (Semicossyphus pulcher) female, and Garibaldi, (Hypsypops rubicundus) in forest of giant kelp (Macrocystis pyrifera) off Santa Barbara Island, California, USA
Smallmouth grunt (Haemulon chrysargyreum) fish swim over a coral reef, Cancun, Mexico. Slow shutter speed image
Pink anemonefish (Amphiprion perideraion) living in symbiotic association with Magnificent sea anemone (Heteractis magnifica), Palau
Translucent coral goby (Bryaninops erythrops) in Seapen (Virgularia gustaviana). Komodo National Park, Indonesia. Indo-West Pacific
Moluccan cardinalfish (Ostorhinchus moluccensis) next to hydroids, brittlestars and sea squirts. Tulamben, Bali, Indonesia
Parrotfish (Scarus / Chlorurus sp. ) sleeping at night, New Caledonia, Pacific OceanParrotfish (Scarus / Chlorurus sp.) sleeping at night, New Caledonia, Pacific Ocean
Ocean Pacifique - Nouvelle caledonie - Gros plan de la tete d un nason licorne (Naso brevirostris) de nuit Pacific Ocean - New Caledonia - A close-up of the head of a spotted or longnose unicornfish
Scalefin anthias (Pseudanthias squamipinnis) male with a long exposure. Ras Katy, Sharm El Shiekh, Sinai, Egypt. Red Sea. Commended in the GDT Photography competition 2019
Yellow pygmy goby (Lubricogobius exiguus) portrait, Kashiwajima, Japan
Canary fish (Epinephelus marginatus) in reef. El Hierro, Canary Islands
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Atlantic salmon (Salmo salar) on breeding territory in the River Ness, Scotland, UK, January
Big Gulf grouper (Mycteroperca jordani), Cabo Pulmo Marine National Park, Baja California Sur, Mexico
Humped rockcod (Gobionotothen gibberiformis), Antarctic Peninsula, Antarctica
Ring-tailed / Golden cardinalfish (Ostorhinchus aureus) male incubating its eggs in its mouth, Sulu sea, Philippines
Stargazer (Uranoscopus sulphureus) half buried in the sand, Sulu sea, Philippines
Grayling (Thymallus thymallus) in the upper reaches of the Lena River. Baikalo-Lensky Reserve, Siberia, Russia. August 2018
Forsters hawkfish (Paracirrhites forsteri) on a coral reef. Jackson Reef, Tiran, Sinai, Egypt. Strait of Tiran, Gulf of Aqaba, Egypt
Reef manta (Mobula alfredi) female swimming close to a coral reef, while cleaner wrasse (Labroides dimidiatus), tiny by comparison, pick parasites from her lips
Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island
Midnight snapper (Macolor macularis) shelters in a cavern on a coral reef with white soft corals (Scleronephthya sp.) and solderfish. Vavuu Atoll, Maldives. Indian Ocean
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef
Whitemargin stargazer (Uranoscopus sulphureus) sets an ambush in the sand at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Atlantic salmon (Salmon salar) under river ice. Quebec, Canada. October
Banggai cardinalfish (Pterapogon kauderni) with a Corkscrew or Long tentacle anemone (Macrodactyla doreensis). Lembeh Strait, North Sulawesi, Indonesia
Potrait of a Cuckoo wrasse (Labrus bimaculatus) Loch Linnhe, west coast of Scotland, UK, September
Cuckoo wrasse (Labrus mixtus) swims through a kelp forest (Laminaria sp) Porthkerris, Lizard Peninsular, Cornwall, UK August
Grayling (Thymallus) under Ice, Lake Baikal, Siberia, Russia
Tompot blenny (Parablennius gattorugine) male peers out of its den, in leg of Swanage Pier, Swanage, Dorset, UK, English Channel, August
Brandts cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus), beneath an oil rig
Two male Dusky groupers (Epinephelus marginatus) fighting for territory during mating season at baixa do meros, a small pinnacle at Formigas islets Protected area, near Santa Maria Island, Azores
Wolf fish (Anarhichas lupus) portrait, Akuyeri, Iceland. North Atlantic Ocean
Arctic grayling (Thymallus arcticus) displaying his vibrant dorsal fin. North Park, Colorado, USA
False clown anemonefish (Amphiprion oceallaris) guarding their eggs laid on a coral rock beneath a sea anemonePhilippines. Bohol Sea, Tropical West Pacific Ocean
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
Gorgonian sea fans (Melithaea sp. ) with a large school of Pygmy sweepers (ParapriacanthusGorgonian sea fans (Melithaea sp.) with a large school of Pygmy sweepers (Parapriacanthus ransonetti) Andaman Sea, Thailand
Red snappers / Two-spot red snappers (Lutjanus bohar) mass shoal of fish spawning, Palau, Philippine Sea. A few black and white snappers (Macolor niger) have crawled into the school to eat the eggs
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
Crevalle Jack (Caranx hippos) and other fish entering Homosassa Spring for the winter, Homosassa Springs Wildlife Park, Florida, USA. March. Photographed for the Freshwater Project
School of Bonito (Sarda orientalis) in open water. Darwin Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Bigeye trevally (Caranx sexfasciatus) forming a circular shoal in blue water, on the drop off of a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
A threadfin hawkfish (Cirrhitichthys aprinus) on a barrel sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Whitebar surgeonfish (Acanthurus leucopareius) school in undewater cave, oxygenated water brought down by Pacific ocean surge. Kona, The Big Island, Hawaii