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Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Mandarinfish (Synchiropus splendidus) Yap, Micronesia
Bignose unicornfish (Naso vlamingii) profile of male displaying (the blue stripe at the bottom of the frame is the tail of another individual) Baa Atoll, Maldives. Indian Ocean
Blackspotted stingray (Taeniurops meyeni) with a remora (Echeneis naucrates) attached to its belly swimming in the open water with a diver in the background. Ari Atoll, Maldives, Indian Ocean
Galapagos barnacle blenny (Acanthemblemaria castroi) looking out from its home in an old barnacle shell. Cape Marshall, Isabela Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
Bigeye trevally (Caranx sexfasciatus) forming a circular shoal in blue water, on the drop off of a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Black Bream (Spondyliosoma cantharus) washed up on shore, Sark, British Channel Islands, August
RF - Orange skunk clownfish (Amphiprion sandaracinos) in anemone, Puerto Galera, Philippines
Big-eye or Goggle-eye (Priacanthus hamrur) and Crown squirrelfish (Sargocentron diadema) on wreck colonised by corals, Red Sea, Egypt. January
Kaleidoscopic image of coral reef with soft corals (Dendropnethya) shoal of Yellow back fusilliers (Caesio teres) West Papua, Indonesia
Spinecheek anemonefish (Premnas biaculeatus) male with bleached anemone, Bunaken National Park, North Sulawesi, Indonesia
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
Emperor angelfish (Pomacanthus imperator) swimming past coral reef. Egypt, Red Sea
Yellowbar angelfish (Pomacanthus maculosus) Egypt, Red Sea
Gorgonian sea fan (Melithaea sp. ) with Lyretail anthias (Pseudanthias sqamipinnis)Gorgonian sea fan (Melithaea sp.) with Lyretail anthias (Pseudanthias sqamipinnis). Similan Islands, Andaman Sea, Thailand
RF - Shoal of Slender pinjalo snappers (Pinjalo lewisi) swimming over soft corals on coral reef. West Papua, Indonesia
Ring-tailed Cardinalfish (Apogon aureus) West Papua, Indonesia
Crown spotty blenny (Laiphognathus longispinis) Pak Lap Tsai, Sai Kung East Country Park, Hong Kong, China
Blue shark (Prionace glauca) with Pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
Atlantic salmon (Salmo salar) jumping up waterfall during spawning migration upstream, Ume River, Sweden. July. Photographed for The Freshwater Project
Painted cardinalfish (Archamia fucata) Yap, Micronesia
European barracuda (Sphyraena sphyraena) and Bluefish (Pomatomus saltatrix) circling baitball of Atlantic horse mackerel (Trachurus trachurus) Formigas Islets, Azores
Bullethead parrotfish (Chlorurus sordidus) male, close up of eye, whilst sleeping at night. The Alternatives, Sinai, Egypt. Red Sea
Whitebelly damselfish (Amblyglyphidodon leucogaster) pair spawning / laying eggs on a seafan (Annella mollis). Ras Mohammed National Park, Sinai, Egypt. Red Sea
Goby (Bryaninops sp) on coral branch, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Reef and shoal of Jewel fairy basslet (Pseudanthias squamipinnis) South Point dive site, Sanganeb reef, Sudan, Red Sea
Underwater view of Brown bear (Ursus arctos) paw fishing for Sockeye salmon (Oncorhynchus nerka) with paw outstretched. Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August
Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp)Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp.) diving towards coral reef to avoid predators. North Ari Atoll, Maldives. Indian Ocean
Oriental sweetlips (Plectorhinchus vittatus) pair. South Ari Atoll, Madlives. Indian Ocean
Reticulate stargazer (Uranoscopus sp. ) portrait. Ambon Bay, Ambon, Maluku Archipelago, IndonesiaReticulate stargazer (Uranoscopus sp.) portrait. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Bluestreak cleaner wrasse (Labroides dimidiatus) just swimming out of mouth of Oblique / Diagonal banded sweetlips (Plectorhinchus lineatus) whilst cleaning / feeding, New Caledonia. Pacific Ocean
Detail of the pectoral fin of Roundhead parrotfish (Scarus viridifucatus) coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
School of Bluespine unicornfish (Naso unicornis) gathered during the summer breeding season. Yolanda Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
School of young European perch (Perca fluviatilis) in Altausseer lake, Austria, July
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
A signal blenny (Emblemaria hypacanthus) displays its showy dorsal fin as it extends out of its tube home. La Paz, Mexico. Sea Of Cortez, East Pacific Ocean
Pink anemonefish (Amphiprion perideraion) in Purple magnificent sea anemone (Heteractis magnifica) Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Reticulate stargazer (Uranoscopus sp. ) emerging from the sandReticulate stargazer (Uranoscopus sp.) emerging from the sand. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
