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Atlantic salmon (Salmo salar) moving upriver to spawn. Lligwy River nr Betws y Coed, Gwynedd, North Wales, UK, November
Apollo butterfly (Parnassius apollo) on thistle head in alpine meadow. Nordtirol, Austrian Alps
Brown Hare (Lepus europaeus) portrait gazing alertly. Perthshire, Scotland, June
Domestic dog, Scottish Terrier / Aberdeen Terrier, studio portrait
Brown bear (Ursus arctos) running through water, fishing, Kamchatka, Far east Russia, August
Brown bear (Ursus arctos) walking beside lake, Kamchatka, Far east Russia, August
Brown bear (Ursus arctos) standing beside lake looking for salmon, Kamchatka, Far east Russia, August
South African fur seal (Arctocephalus pusillus pusillus) swimming near surface, Seal Island. False Bay, South Africa, July 2010
Hawaiian goose / Nene (Branta sandvicensis) lookig down, Slimbridge, Gloucestershire, UK, Captive, native to Hawaii, USA, Vulnerable species, September
Zanzibar Red Colobus (Procolobus kirkii) head portrait, Jozani Chwaka Bay NP, Zanzibar, Tanzania
Proboscis Monkey (Nasalis larvatus) female sitting in tree, partially obscured by soft focus leaves in foreground, Bako National Park, Sarawak, Borneo, Malaysia
Australian Shepherd, blue-merle, head portrait with autumn leaves behind
Norfolk Terrier portrait sitting in autumn foliage
Red deer (Cervus elaphus) stag amongst winter vegetation, Salburua Park, Alava, Spain, November
African lion (Panthero leo) male giving flehmen grimace, Tanzania, East Africa
Little owl (Athene noctua) perched on post at dusk, Carmarthenshire, Wales, UK, Captive
Polar bear (Ursus maritimus) portrait of curious young male, covered in snow from rolling around / playing. On a barrier island during autumn freeze up, Bernard Spit
Ptarmigan (Lagopus mutus) male in flight over snow covered Cairngorm Mountains, Scotland, February. (non-ex)
Old male African lion (Panthera leo) portrait, Sabi Sand Game Reserve, South Africa, June
Catesbys snail-eating snake (Dipsas catesbyi) on Heliconia plant, Pacaya Samiria National Reserve, Amazon Basin, Loreto, Peru
Golden eagle (Aquila chrysaetos) in flight over snow, Alps, Austria, controlled conditions
Panther Chameleon {Furcifer pardalis} walking along branch on coast, Masoala Peninsula National Park, north east Madagascar
Giraffe necked weevil {Trachelophorus giraffa} male on leaf of host plant (Dichaetanthera cordifolia) tropical rainforest, Mantadia NP, Madagascar
An adult female California sea lion (Zalophus californianus) head portrait, in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Eye and spiracle detail of a Bluespotted / Ribbontail Stingray (Taeniura lymma) Sipadan Island, Sabah, Borneo, Malaysia. May
California sea lion pup (Zalophus californianus) portrait in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Atlantic grey seal (Halichoerus grypus) swimming beneath the surface, Lundy Island, Devon, England, UK. July 2010. Highly commended, Coast and Marine category
Lemon shark (Negaprion brevirostris) in shallow water with reflection at the surface. Little Bahama Bank, Bahamas
Polar bear (Ursus maritimus) head close-up portrait of an adult male, with snowflakes on fur, Barter Island, 1002 area of the Arctic National Wildlife Refuge, Alaska
Red Kite (Milvus milvus) diving towards prey whilst in flight, Wales, UK, March
Crested Newt (Triturus cristatus carnifex) underwater view of female swimming on surface, Traversari lake, Campigna National Park, Emilia Romagna and Tuscany regions, Italy
Sri Lankan Leopard (Panthera pardus kotiya) soft focus running, captive
As soon as it entered the water the young Olive ridley sea turtle (Lepidochelys olivacea) struggles against the swell to swim away from the coast
Mountain hare (Lepus timidus) running across wind swept snow field, near Carrbridge, Cairngorms National Park, Scotland, January 2010
Kromfohrlander running / racing in field towards camera
Small Munsterlander head portrait
Dalmatian, head portrait, male aged 4 years
Alaskan Malamute portrait of blue coated adult
Fieldfare (Turdus pilaris) feeding on Rowan berries (Sorbus hupehensis) in winter, Perthshire, Scotland. WINNER: Young Wildlife Photographer of the Year 2010
Adult male Bornean orangutan (Pongo pygmaeus) in rainforest canopy, Gunung Palung National Park, Borneo, West Kalimantan, Indonesia
Arctic tern (Sterna paradisaea) in flight, vocalising, aggresive behaviour, Farne Islands, Northumberland, UK, June
Arctic tern (Sterna paradisaea) in flight, calling, Farne Islands, Northumberland, UK, June
Cheetah (Acinonyx jubatus) running at full speed chasing Springbok prey, Kalahari Desert, Southern Africa
White-spotted leaf frog (Cochranella albomaculata) captive, from Central America. Please credit Michael Kern / Zoo Atlanta
Checkered Garter Snake (Thamnophis marcianus marcianus) coiled with tongue protruding. Fennessey Ranch, Refugio, Corpus Christi, Coastal Bend, Texas Coast, USA
Jaguar (Panthera onca) female, hiding in dense high grass. Cuiaba river, Pantanal, Mato Grosso do Sul Province. Brazil, South America
Portrait of German Shorthair Pointer in thicket of pink berries, Illinois, USA
Head portrait of black and tan smooth coated Dachshund with zinnias, Illinois, USA