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Atlantic Puffin (Fratercula arctica) resting on a snow bank, Isle of Hornoya, Varanger, Norway, April. Bookplate from Danny Greens The Long Journey North
Fulmer (Fulmras glacialis) in flight near blue glacier, Svalbard, July. Bookplate from Danny Greens The Long Journey North
White tailed Sea Eagle (Haliaeetus albicilla) in flight, Norway, July. Bookplate from Danny Greens The Long Journey North
Tawny Owl (Strix aluco) adult female hiding behind tree trunk, trained bird, Somerset, UK, January
Split image of Blue shark (Prionace glauca) close to the surface, Pico Island, Azores, Portugal, Atlantic Ocean, July
Detail of magnificent Sea anemone (Heteractis magnifica) Maldives, Indian Ocean
Lammergeier (Gypaetus barbatus) close up portrait, Spain, November
Capercaillie (Tetrao urogallus) in flight, Kuusamo, Finland, January
Jogi, a Jack Russell Terrier cross breed, male playing in the snow. Germany
Eurasian Lynx (Lynx lynx) adult sitting on boulder covered in deep snow in beech (Fagus sylvatica forest, captive in enclosure of the Bavarian Forest National Park, Germany, February
Harvest mouse (Micromys minutus) on barley cereal, Yorkshire, UK Captive
Andean condor (Vultur gryphus) in flight over mountain peaks, Torres Del Paine National park, Chile
Humpback Whale (Megaptera novaeangliae) calf, Tonga, South Pacific, September
Portrait of a Yellow seahorse (Hippocampus guttulatus), Capo Galera, Alghero, Sardinia, Italy, Mediterranean Sea
Horsefly (Philipomyia aprica) Aosta Valley, Monte Rosa Massif, Pennine Alps, Italy. July
Mountain Clouded Yellow (Colias phicomone) profile portrait, Aosta Valley, Monte Rosa Massif, Pennine Alps, Italy
Golden Eagle (Aquila chrysaetos) portrait in snow, Utajarvi, Finland February
Arctic Tern (Sterna paradisaea) in flight, silhouetted against the moon at dusk, Uto Finland July
Zanzibar red colobus (Piliocolobus kirkii) in tree, Jozany Forest, Zanzibar, Tanzania. Highly endangered
Polar Bear (Ursus maritimus) head portrait with blue tongue sticking out, Svalbard, Norway
Male Marsh pride lion (Panthera leo) on a moonless night, Masai Mara, Kenya. Taken with infra red camera, September
Snow Goose (Chen caerulescens), coming in to land. Bosque del Apache, New Mexico, USA, November
Marine Iguana (Amblyrhynchus cristatus) on rocks with wave splash. Galapagos, Ecuador
Red Squirrel (Sciurus vulgaris) adult reaching up. Cairngorms National Park, Scotland, UK, March
Polar bear (Ursus maritimus) adult walking over newly formed pack ice during fall freeze up, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska, USA
Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA
Spectacled caiman (Caiman yacare) walking on a sandy shore at the edge of a river at night, Pantanal, Brazil
Little egret (Egretta garzetta) attempting to catch fish, Camargue, France, July
Whooper swan (Cygnus cygnus) in snowfall. Martin Mere Wetlands Trust, Lancashire, UK, January. Animal Portraits category, British Wildlife Photographer Awards (BWPA) competition 2012
Purple gallinule (Porphyrio martinicus) moving across waterlily covered surface, Everglades National Park, Florida, USA, March
Big fin reef squid (Sepioteuthis lessoniana) descends from open water to the reef at night. Arafur Sea, Triton Bay, West Papua, Indonesia
Whiskered auklet (Aethia pygmaea), Iony Island / Jonas Island, Sea of Okhotsk, Far East Russia, July
Robin (Erithacus rubecula) perched on rosehip branch during snowfall. UK, December
Black faced Impala (Aepyceros melamis petersi) female jumping, Etosha National Park, Namibia
Malachite kingfisher (Alcedo cristata) perched on rush, Intaka Island, Cape Town, South Africa, February
Close-up of Brown bear (Ursus arctos) in lake, Kamchatka, Far east Russia, August
Highland bull (Bos taurus) face close up in freezing fog, Berwickshire Scotland, December
Brown trout (Salmo trutta) resting on river bed near the Aigas Field Studies Centre, Inverness-shire, Scotland, July
Clouded leopard (Neofelis nebulosa) Guwahati, Assam, India, captive
Mistle Thrush (Turdus viscivorus) in snow. Hertfordshire, England, UK, February
Southern plains grey langur ( Semnopithecus entellus / Presbytis entellus) close up face profile, Sawai Modhopur, Rajasthan, India
Boxer looking over closed door, standing on hind legs
Spotted scorpionfish (Scorpaena plumieri) lying motionless and camouflaged in soft corals. West Palm Beach, Gulf Stream, West Atlantic Ocean, Florida, USA
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Yellow goatfish (Mulloidichthys martinicus) swimming above the reef, West End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Spotted owl (Strix nebulosa) Gifford-Pinchot National Forest, Washington USA
Grey Heron (Ardea cinerea) taking a fish from water. Kiskunsagi National Park, Hungary, May
Freerange Gloucester old spot domestic pig (Sus scrofa domestica) portrait with ears covering eyes, UK