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Boxer looking over closed door, standing on hind legs
Spotted scorpionfish (Scorpaena plumieri) lying motionless and camouflaged in soft corals. West Palm Beach, Gulf Stream, West Atlantic Ocean, Florida, USA
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Yellow goatfish (Mulloidichthys martinicus) swimming above the reef, West End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Spotted owl (Strix nebulosa) Gifford-Pinchot National Forest, Washington USA
Grey Heron (Ardea cinerea) taking a fish from water. Kiskunsagi National Park, Hungary, May
Freerange Gloucester old spot domestic pig (Sus scrofa domestica) portrait with ears covering eyes, UK
Cotswolds Lion rare breed sheep (Ovis aries) and the village of Naunton at sunset, Gloucestershire, UK, June 2001
Indian elephant (Elephas maximus) close up of skin and tail, captive
Red river hog (Potamochoerus porcus) close up of head, captive
Red Grouse (Lagopus lagopus scoticus) in Heather. Peak District, UK, August
Harlequin Ladybird (Harmonia axyridis) melanic form, on turning leaf. Sheffield, October
Red fox (Vulpes vulpes) head portrait, The Netherlands, March
Wild cat (Felis silvestris) in snowstorm captive, Germany
Grizzly bear (Ursus arctos horribilis) walking, Yellowstone National Park, Wyoming, USA, October
Blue Shark (Prionace glauca) at the sea surface. Santa Maria, Azores, August
Mute swan (Cygnus olor) on water, Wheatfen, Norfolk, UK, November
Heath fritillary (Melitaea athalia) resting on scabious flower, Nordtirol, Austrian Alps, Austria, August
Jumping spider (Euophrys frontalis) male amongst flower petals, Peak District National Park, Derbyshire, UK, June
Harvest mouse (Micromys minutus soricinus) feeding on blackberry, captive, Dorset, UK, October
German Hound male, 6 months
Sparkling violetear hummingbird (Colibri coruscans) at base of Chimborazo Volcano (Highest mountain in Ecuador), Andes, Ecuador, South America
Brown Long-eared bat (Plecotus auritus) drinking from a lily pond, Surrey, UK. Overall WINNER of the Audubon Society of Greater Denver Share the Viewae competition 2011
Male Indian peafowl (Pavo cristatus) displaying, captive
Black swan (Cygnus atratus) captive
Comtois Horse (Equus caballus). Camargue reserve, Rhone, France, September
Male Whinchat (Saxicola rubetra) perched on Oil seed rape (Brassica napus) in a field, Vosges, France, April
Roe deer (Capreolus capreolus) at sunset, Vosges, France, August
Tropical Butterfly (Adepha) on a leaf with wings spread. Danum Valley, Sabah, Borneo, Malaysia
Ichneumon wasp (Compsocryptus sp. ?) in flight, Williamson county, Texas, USA, NovemberIchneumon wasp (Compsocryptus sp.?) in flight, Williamson county, Texas, USA, November
Bridled nailtail wallaby (Onychogalea fraenata) Queensland, Australia, Endangered species, October
Dipper (Cinclus cinclus) perched on moss-covered waterfall, Peak District NP, Derbyshire, UK, May, Highly Commended, Habitat category, British Wildlife Phototgraphy Awards (BWPA) competition 2011
Southern elephant seal (Mirounga leonina) close up of sleeping individual, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Marbled White Butterfly (Melanargia galathea) resting on grass stem, Devon, UK July. Winner of Seasons category of the BWPA 2011 competition
Great Blue Shark (Prionace glauca), dorsal view against dark water. Over 20 million Blue Sharks are fished annually, as bycatch and due to the high value of their fins
Brown hare (Lepus europaeus) running past hay barn in snow storm, Lapua, Finland, March. WINNER, Animals in their Environment category, 2011 WILDLIFE PHOTOGRAPHER OF THE YEAR competition
Common green tree frog (Litoria caerulea) in rainforest, Iron Range National Park, Cape York Peninsula, North Queensland, Australia, October
A rare Sorraia stallion cantering, Coudelaria Nacional (National Stud Farm), in Alter do Chao, District of Portalegre, Alentejo, Portugal
Common Swallowtail butterfly (Papilio machaon) resting on Meadow Clary (Salvia pratensis) flowers, on a white background. Nordtirol, Tirol, Austrian Alps, Austria, 1700 metres altitude, July
Beagle hound in wild purple Violets, Arcadia, Wisconsin, USA
Red squirrel (Sciurus vulgaris) standing on its hind legs, Cairngorms National Park, Scotland, UK, March. Non-ex
Grey seal (Halichoerus grypus) adult female among breaking waves, Lincolnshire, UK Non-ex
Arctic tern (Sterna paradisaea) adult hovering directly in front of the sun, defending nest from perceived threat, Farne Islands, UK. June Non-ex
Ural Owl (Strix uralensis) perched. Bieszczady National Park, the Carpathians, Poland, December
Blue Bearded Collie puppy, 3 months, yawning
Border Collie bitch, running towards the camera
Chinchilla Persian female cat, 6 years
Yellow Labrador Retriever puppy, 7 weeks