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Arctic Tern (Sterna paradisaea) in flight, silhouetted against the moon at dusk, Uto Finland July
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November
Heermanns Gulls (Larus heermanni) in flight. Ensenada, Baja California Peninsula, Mexico, December
Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA
Emperor angelfish (Pomacanthus imperator) with soft corals and sweepers. Egypt, Red Sea
Killer WHale / Orca (Orcinus orca) bull and cow in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May
Purple gallinule (Porphyrio martinicus) moving across waterlily covered surface, Everglades National Park, Florida, USA, March
Flock of Greater flamingo (Phoenicopterus ruber) in flight, Camargue, Southern France, November 2006
Cowboys rounding up a band of Criollo pure pedigree mares and foals, Estancia Ita Maria, Misiones, Paraguay, January 2012
Aggregation / bloom / swarm of Moon Jellyfish (Aurelia aurita) in the clear waters of Loch Na Keal, off the Isle of Mull, north west Scotland, UK, June, Photographed through the surface
Grizzly bear (Ursus arctos horribilis) walking, Yellowstone National Park, Wyoming, USA, October
Manta Ray (Manta Birostris) and Devil Ray (Mobula tarapacana) in background, accompanied by Pilot Fish (Naucrates ductor) and Remora. Santa Maria Island, Azores, September
Blue Shark (Prionace glauca) at the sea surface. Santa Maria, Azores, August
Mute swan (Cygnus olor) on water, Wheatfen, Norfolk, UK, November
Flock of Cape petrels (Daption capense capense) in flight against the sea. Against the sun. South Shetland Islands, Antarctica, January
Caribbean reef shark (Carcharhinus perezi) Jardines de la Reina National Park, Cuba
Jersey tiger moth (Euplagia quadripunctaria) large numbers flying to and resting on Sweetgum trees (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011
Acacia sp. at night, with a herd of Plains Zebra (Equus quagga) beneath. Near Ndutu, Nogorongoro Conservation Area, Serengeti National Park, Tanzania
Brown Long-eared bat (Plecotus auritus) drinking from a lily pond, Surrey, UK. Overall WINNER of the Audubon Society of Greater Denver Share the Viewae competition 2011
Common dolphins ( Delphinus delphis) swimming below the surface, Atlantic Ocean, Portugal, October
Two Scarlet macaws (Ara macao) flying, captive
Ichneumon wasp (Compsocryptus sp. ?) in flight, Williamson county, Texas, USA, NovemberIchneumon wasp (Compsocryptus sp.?) in flight, Williamson county, Texas, USA, November
Great Blue Shark (Prionace glauca), dorsal view against dark water. Over 20 million Blue Sharks are fished annually, as bycatch and due to the high value of their fins
A rare Sorraia stallion cantering, Coudelaria Nacional (National Stud Farm), in Alter do Chao, District of Portalegre, Alentejo, Portugal
Termites (Isoptera) flying upwards towards a street lamp, India
Wild / feral Dulmen ponies (Equus caballus) close up of herd of mares and foals running on the Duke of Croys estate, Meerfelder Bruch, North Rhine-Westphalia, Germany, May 2011
Jackdaws (Corvus monedula) abstract view of flock in flight, Derbyshire, UK, December. Non-ex
Hand reared Whooper Swans (Cygnus cygnus) in flight. Birds imprinted and trained by Lloyd and Rose Buck for filming purposes. Somerset, UK, January 2011
Black Rhinoceros (Diceros bicornis) charging. Etosha National Park, Namibia, January
A male Leafy Seadragon (Phycodurus eques) swimming. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
A juvenile Weedy Seadragon (Phyllopteryx taeniolatus) in its habitat. Flinders Jetty, Melbourne, Victoria, Australia, March
Long Water Measurer (Hydrometra stagnorum) drinking by piercing surface tension with mouthparts. Captive, UK, July
Alpine Chough / Yellow-billed Chough (Pyrrhocorax graculus) flock in flight against mountains, Dolomites, Italy. Digital composite
Red Fox (Vulpes vulpes) pouncing in grass. Vosges, France, July
Roe Deer (Capreolus capreolus) swimming across River Allier, France, February
Chestnut Paso peruvian stallion running, California, USA
Two Common Bottlenose Dolphin (Tursiops truncatus) in shallow water. Punta Allen, Sian Ka an Biosphere Reserve, Quintana Roo, Yucatan Peninsula, Mexico
Golden eagle (Aquila chrysaetos) about to land on snow, Oulanka NP, Finland, February
Grey Heron (Ardea cinerea) leaving footprints while walking through snow. The Netherlands, January
South African fur seal (Arctocephalus pusillus pusillus) swimming near surface, Seal Island. False Bay, South Africa, July 2010
Atlantic grey seal (Halichoerus grypus) swimming beneath the surface, Lundy Island, Devon, England, UK. July 2010. Highly commended, Coast and Marine category
Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean
Black browed albatross (Thalassarche melanophrys) flying in 60 knot wind at Cape Horn, Chile
Bottlenose dolphins (Tursiops truncatus) in shallow water over a sand bank. Sandy Ridge, Little Bahama Bank, Bahamas. Tropical West Atlantic
Common / Eurasian cranes (Grus grus) small flock of juveniles taking flight over the frozen, snow covered pastureland on a foggy winter morning
Red Kite (Milvus milvus) diving towards prey whilst in flight, Wales, UK, March
Sri Lankan Leopard (Panthera pardus kotiya) soft focus running, captive
Mountain hare (Lepus timidus) running across wind swept snow field, near Carrbridge, Cairngorms National Park, Scotland, January 2010