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Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Leatherback Turtle (Dermochelys coriacea) hatchling crossing a beach to get to the sea. Cayenne, French Guiana, July
Chilean flamingos (Phoenicopterus chilensis) in flight over mountain peaks with glacier in the distance, Torres Del Paine National Park, Chile
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Nudibranch (Janolus cristatus) Vela Luka, Korcula Island, Croatia, Adriatic Sea, Mediterranean
Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, Habitat division, British Wildlife Photography Awards (BWPA) competition 2012
Head portrait of grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
European hare (Lepus europaeus), Wirral, England, UK, May
African elephant (Loxodonta africana) herd walking in the plains during the dry season, Masai-Mara Game Reserve, Kenya. October
Common toad (Bufo bufo) tadpole swimming seen from below, Belgium, June
Line of Leaf-cutter ants (Atta sp) carrying leaves, digital composite
Portrait of a grey Andalusian stallion, in Ciutadella, Menorca, Spain
Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009
RF- Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
King Eider (Somateria spectabilis) flying low over water. Floe edge, Arctic Bay, Nunavut, Canada, April
Dromedary camel (Camelus dromedarius) train crossing Merzouga dunes, Tafilalt, Sahara, Morocco, March 2007
Green Turtle, (Chelonia mydas), Swimming over volcanic sandy bottom, Armenime cove, South Tenerife coast, Canary Islands, Spain, Atlantic Ocean, May 2011
Gentoo penguins (Pygoscelis papua) group walking along snow, Cuverville Island. Antarctic Peninsula, Antarctica
Caribbean reef squid (Sepioteuthis sepioidea) swimming in open water at night, Guadeloupe Island, Mexico. Caribbean
Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Snow Leopard (Panthera uncia) climbing up mountain slope, Hemis National Park, India, February
RF - Wild pinto Mustang stallion running in snow, Black Hills Wild Horse Sanctuary, South Dakota, USA. January
Green Sea Turtle (Chelonia mydas) returning to sea after Egg laying, Raine Island, Australia. Raine Island is the largest and most important green sea turtle nesting area in the world, with over 14
Group of California sea lions (Zalophus californianus) in a kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Canada goose (Branta canadensis) adult swimming leading five goslings, Ithaca, New York, USA
Juvenile male Peregrine falcon (Falco peregrinus) in flight chasing his parent who has Feral pigeon (Columba livia) kill in claws, Bristol, England, UK, June
Female Capybara (Hydrochoerus hydrochaeris) swimming in line with young after escaping a Jaguar attack (Panthera onca palustris) in a lagoon off the Paraguay River
Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Tibetan Wild Ass / Kiang, (Equus kiang) running, ChangThang, Tso Kar lake, at altitude of 4600m, Ladakh, India
Chamois (Rupicapra rupicapra) running down ridge, in early morning light. Lausons Valley, Gran Paradiso National Park, Italy, September
Snowy owl (Bubo scandiacus) in flight, Boundary Bay, British Columbia, Canada. February. First place in the Overall Category of the 2013 Great Backyard Bird Count photo competition
Golden Eagle (Aquila chrysaetos) in flight, snowing, Utajarvi, Finland, February
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Flatanger, Norway, June
Brown hare (Lepus europaeus) adult running in wintery landscape. Derbyshire, UK, January
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
Southern stingray (Hypanus americanus) female swimming over a shallow sand bank, under morning sun, North Sound, Grand Cayman, Cayman Islands. British West Indies, Caribbean Sea
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Cape buffalo (Syncerus caffer) crossing the Chobe River, followed by swarm of flies, Chobe Game Reserve, Botswana, Africa
Wildebeest (Connchaetes taurinus) running down bank of the Mara River during migration, Masai Mara, Kenya
People trekking with Dromedary camels (Camelus dromedarius) through chalk rock formations created by sandstorms, White desert, Sahara, Egypt, February 2009. No release available
Blue shark (Prionace glauca) Pico Island, Azores, Portugal, Atlantic Ocean, July
Horse in snow storm with shed in background, USA
Dall sheep (Ovis dalli) yearling lamb running across alpine tundra, Denali National Park, Alaska, USA
Whale Shark (Rhincodon typus) accompanied by Pilot Fish (Naucrates ductor) and Remora. The biggest fish in the world has only started to visit the Azores in the past three years due to water
