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Stellers Eider (Polysticta stelleri) male swimming, Varangerfjord, Norway, April, Vulnerable species. Bookplate from Danny Greens The Long Journey North
Great Grey owl (Strix nebulosa) taking off from the ground, Finland, April. Bookplate from Danny Greens The Long Journey North
White-tailed Sea Eagle (Haliaeetus albicilla) being attacked by a Common Gull (Larus canus) in flight. Norway, July. Magic Moments book plate, page 51
Mountain hare (Lepus timidus) in white winter coat stretching - in snowy habitat, Scotland. Highly Commended in the Mammals category of the GDT European Photographer of the Year Competition 2015
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
Common / Eurasian crane (Grus grus) flock in flight with snow topped Pyrenees mountains in the background, during southward migration period, Haute Pyrenees, Gascony, France. December 2009
Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Herd of Tibetan Wild Asses / Kiang (Equus kiang) ChangThang, Tso Kar lake, at altitude of 4600m, Ladakh, India, October 2012
Massing herds of White bearded wildebeest (Connochaetes taurinus albojubatus) on migration, Serengeti National Park, Tanzania
Nile crocodile (Crocodylus niloticus) making its way through sand to the Rufiji River, Selous Game Reserve, Tanzania
Wildebeest (Connochaetes taurinus) crossing the Mara river, Masai-Mara Game Reserve, Kenya
Sanderling (Calidris alba) running on the tideline, Snettisham, Norfolk, December. Bookplate from Danny Greens The Long Journey North
Common Raven (Corvus corax) two in flight during snow storm, Kuusamo, Finland, April
Wild Cat (Felis silvestris). Vosges, France, August
Wild yak (Bos mutus) running, Kekexili, Qinghai, Tibetan plateau, China, December
Siskins (Carduelis spinus) fighting over food. Oisterwijk, The Netherlands, March
Blacktip reef sharks (Carcharhinus melanopterus) swimming in shallow crystal clear water, Aldabra Atoll, Seychelles, Indian Ocean
Bullers shearwater (Puffinus bulleri) in flight low over the water, showing upperwing. Off Whitianga, Coromandel Peninsula, New Zealand
White horses of the Camargue, herd running through water, Camargue, Southern France
A rare East Bulgarian stallion cantering at the Kabiuk National Stud, Shumen, Bulgaria, February 2012
A flock of Greylag Goose (Anser anser) flying in formation above misty fields. The Netherlands, March
Western grebe (Aechmophorus occidentalis) pair swimming, with reflections, Bolsa Chica Ecological Reserve, California, USA, February
Leatherback Turtle (Dermochelys coriacea) returning to the sea after egg laying. Cayenne, French Guiana, June
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
Great Horned owl, (Bubo virginianus) in flight, Regina, Saskatchewan, Canada, March
Leaf cutter ant (Atta sp) female worker carrying leaf to nest, with a smaller minor ant riding. The minors job is to protect her sister from parasitic flies. Costa Rica, December
Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November
Little Auks (Alle alle) flying low above surface in front of iceberg. Spitsbergen, Svalbard, Norway, July 2010. Highly commended in the Birds category in the GDT competition 2011
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Hoopoe (Upupa epops) in flight, Hungary. Multi flash, long exposure. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition
Emperor penguins (Aptenodytes forsteri) males dive through openings in the sea ice to take their first swim in over 4 months as they have spent the bleak winter incubating their eggs
Puffins (Fratercula arctica) pair take off together to go fishing. Farne Islands, Northumberland, UK, May. Non-ex
A shoal of Ornate Ghost Pipefish (Solenostomus paradoxus) below a black coral bush (Antipathes sp)A shoal of Ornate Ghost Pipefish (Solenostomus paradoxus) below a black coral bush (Antipathes sp.). Misool, Raja Ampat, West Papua, Indonesia, November
Grey Andalusian / Spanish stallion running, California, USA
Barrel Sponge (Xestospongia testudinaria) on coral reef with soft corals (Scleronephthya sp), gorgonian and Lyretail anthias (Pseudanthias squamipinnis). Komodo National Park, Indonesia
Great Tit (Parus major) in flight. Bavaria, Germany, December
European Otter (Lutra lutra) walking across snow while sniffing the ground. The Netherlands, December. Captive
Tanzanian Giant Millipede {Archispirostreptus gigas} Captive, originating from Africa
Silhouette of Brown bear (Ursus arctos) beside Lake Kuril at dawn, Kamchatka, Far east Russia, August 2008
Narwhal (Monodon monoceros) swimming near the surface, seen from the air. Pond Inlet, Canadian Arctic
A pair of Toco Toucans (Ramphastos toco) flying across the Piquiri River at dawn, northern Pantanal, Mato Grosso, Brazil, September. (Digitally Modified)
Burchells Zebra (Equus quagga burchellii) herd walking towards waterhole in the late afternoon, kicking up dust as they come. Etosha National Park, Namibia, Southern Africa
