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Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea
California sea lions (Zalophus californianus) group playing in the sun in the early morning. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
RF - Long exposure of a school of yellowtail surgeonfish (Prionurus punctatus), these fish school to help them access the best feeding areas, which are controlled by aggressive damselfish
RF - Split level photo of a Whale shark (Rhincodon typus) feeding at the surface. Isla Mujeres, Yucatan Mexico. Caribbean Sea
Aggregation of tiny isopods (Santia sp. ) living on the surface of a Blue sponge (Haliclona sp)Aggregation of tiny isopods (Santia sp.) living on the surface of a Blue sponge (Haliclona sp.), there are two colour varieties here, possibly different species
Southern stingrays (Hypanus americanus) swimming over a sand bar in the early morning. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
RF - Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell on the sandy seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Lionfish (Pterois volitans) portrait, invasive species in the Caribbean. East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Portrait of a male Squarespot anthias (Pseudanthias pleurotaenia). Sangeang Island; Sumbawa, Indonesia. Flores Sea
Studio image of multiple Nudibranch species. Gulen, Norway. North East Atlantic Ocean. Digital composite
School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Four California spiny lobsters (Panulirus interruptus) shelter beneath a boulder in a kelp forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Hairy frogfish (Antennarius striatus) on a sandy seabed. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
Brandts cormorant (Phalacrocorax penicillatus) swimming beneath an oil rig. Eureka Rig, Los Angeles, California, United States of America. North East Pacific Ocean
A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
Great white sharks (Carcharodon carcharias) males beneath the surface. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Bigfin reef squid (Sepioteuthis lessoniana) capturing a pelagic shrimp with long red antennae, in midwater, at night. Anilao, Batangas, Luzon, Philippines
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
RF - Green sea turtle (Chelonia mydas) beneath clouds. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
An Australian Sea Lion (Neophoca cinerea) bathing in the sun at the surface. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Giant Pacific Manta Ray (Manta birostris) portrait with Remora fish (Echeneididae) Roca Partida, Revillagigedos Islands, Mexico. East Pacific Ocean. February
Purple short tentacle mushroom coral (Fungia sp. ) close up of polyps. Lembeh StraitPurple short tentacle mushroom coral (Fungia sp.) close up of polyps. Lembeh Strait, North Sulawesi, Indonesia
Pink anemonefish (Amphiprion perideraion) in a purple Magnificent sea anemone (Heteractis magnifica). Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Gorgonian / Fan coral (Solenocaulon akalyx), close up, West Papua, Indonesia
Polar bear (Ursus maritimus) silhouetted at waters edge at sunset, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, September. Vulnerable species
Starfish close up detail, Blairgowrie, Mornington Penisular, Victoria, Australia. Port Philip Bay, Bass Strait
Coral clam (Pedum spondyloideum) close up, Maldives, Indian Ocean
Yellow tube sponge (Aplysina fistularis) on a coral reef, with the camera rotated during the long exposure. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Kelp gull (Larus dominicanus) on rocky shore with crashing waves at sunset, Bird Island, Lamberts Bay, Western Cape province, South Africa, September 2012
Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean
Flatworm (Maiazoon orsaki) Raja Ampat, Irian Jaya, West Papua, Indonesia, Pacific Ocean
RF- Boulder star coral (Montastrea annularis) showing fluorescent green coloration, photographed under blue light at night, to excite fluorescent pigments in the coral, East End, Grand Cayman
DELETE Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
School of Blackfin barracuda (Sphyraena qenie), Kimbe Bay, West New Britain, Papua New Guinea
Yellow and blueback fusilier (Caesio teres) schooling over coral reef, West Papua, Indonesia
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Sunbeams below the surface of the sea, Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Yellow and blueback fusilier (Caesio teres) shoal, Similan Islands, Andaman Sea, Thailand
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Bigeye trevally or jack (Caranx sexfasciatus) shoal, Sipadan, Malaysia
Sandy beach and bright blue ocean, Fuerteventura, Canary Islands. April 2013
Blue sea star (Linckia laevigata) on coral reef, Fiji, South Pacific
Close up of pattern of adult Emperor angelfish (Pomacanthus imperator) Tulamben, Bali, Indonesia. Java Sea
RF - Spinyhead blenny fish (Acanthemblemaria spinosa) extends out of its hole in a red sponge to snare passing food. Caribbean Sea
Longnose hawkfish (Oxycirrhites typus) hiding in coral, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines