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RF - Black Grouse (Tetrao tetrix) male peforming flutter jump display on lek, Cairngorms National Park, Scotland, UK.May (This image may be licensed either as rights managed or royalty free.)
Nilgiri marten (Martes gwatkinsii) walking on a fallen branch in Mukurthi National Park, Tamil Nadu, India
Rhesus macaque (Macaca mulatta), juveniles playing in tree, Keoladeo NP, Bharatpur, India
Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India
Darter ( Anhinga melanogaster) drying wings, Keoladeo NP, Bharatpur, India
White rhinoceros (Ceratotherium simum) male and female, face to face, Mosi Oa Tunya National Park, Zambia
White rhinoceros (Ceratotherium simum) with a long horn, Marakele National Park, South Africa
Olive-backed sunbird (Nectarinia jugularis) female returning to her nest with nesting material, Cape Hillsborough National Park, Queensland, Australia, September
Long exposure of a Dartmoor stream, Devon, England, UK, August
View towards Higger Tor from Over Owler Tor with heather in full bloom. Abandoned millstone can be seen in the foreground. Peak District National Park, Derbyshire, UK. August 2015
Brimstone butterfly (Gonepteryx rhamni) on Devil s-bit Scabious (Succisa pratensis). Peak District National Park, Derbyshire, UK. August
Harebells (Campanula rotundifolia) growing in disused limestone quarry. Peak District National Park, Derbyshire, UK. August
The Pinnacle Stone, Curbar Edge surrounded by flowering heather. Peak District National Park, Derbyshire, UK. August
High altitude Boeri Lake in old volcano crater, Morne Trois Pitons National Park, Dominica, West Indies, Windward Islands
Old trees in the Syndicate Forest National Park, Dominica, Eastern Caribbean. February 2017
Alpine Ibex (Capra ibex) in its landscape, Valsavarenche, Gran Paradiso national park, Aosta Valley, Italy, May
Red fox (Vulpes vulpes) in deep snow emerging from den, Valsavarenche valley, Gran Paradiso national Park, Aosta Valley, Alps, Italy, December
Alpine marmot (Marmota marmota) carrying grass and other nesting material across snow to its nest while another one is on the lookout, Gran Paradiso National Park, Aosta Valley, Italy, April
Chimpanzee female (Pan troglodytes schweinfurthii) sleeping in a nest built in a tree, Kibale National Park, Uganda, January
Female Uganda kob (Kobus kob thomasi) suckling calf. Murchisson Falls National Park, Uganda. February
Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia
Coalition of two African male lions (Panthera leo) resting on the banks of the Luangwa river, dry season, South Luangwa National Park, Zambia
Hawthorn tree and granite outcrop at Holwell tor, sunrise and mist, Dartmoor National Park, Devon, UK. October 2018
Great Mis Tor covered in snow, Dartmoor National Park, Devon, England, UK, January 2015
Great Staple Tor at dusk, Dartmoor National Park, Devon, England, UK, January
Landscape view of Great Staple Tor, Dartmoor National Park, Devon, England, UK, January 2012
Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January
European otter (Lutra lutra) group with one shaking off water, Kiskunsagi National Park, Hungary, October
European otter (Lutra lutra) shaking off water, Kiskunsagi National Park, Hungary, January
Forested island on Loch Lomond, Loch Lomond and the Trossachs National Park, Scotland, UK
Mountain Gorilla (Goriila beringei) resting, Volcanoes National Park, Virunga Mountains, Rwanda
Cuban emerald hummingbird (Chlorostilbon ricordii) Guanahacabibes Peninsula National Park, Cuba
Bee hummingbird (Mellisuga helenae) feeding from flower, Guanahacabibes Peninsula National Park, Cuba
Mountain gorilla (Gorilla beringei beringei), dominant silverback in forest clearing. Volcanoes National Park, Rwanda
Muskox (Ovibos moschatus) in tundra with mountains in background, Dovrefjell National Park, Norway. September 2018
RF - American Bison (Bison bison) male in snow covered in frost, standing by dead trees. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January
Cuban crocodiles (Crocodylus rhombifer) photographed on a crocodile farm started by Fidel Castro in the 1960s in Cienaga de Zapata National Park to help save the species. Cuba
Pack of wolves (Canis lupus). Yellowstone National Park, Wyoming, USA. January
Freshwater pools on Waterfall Creek, at the top of Gunlom Falls. Mary River, Kakadu National Park, Northern Territory, Australia. March 2014
Bamboo lemur (Hapalmur griseus) sitting on tree stump. Andasibe-Mantadia National Park, Madagascar
Thomsons gazelle (Eudorcas thomsonii) herd. Serengeti National Park, Serengeti, Tanzania
Beech woodland near Blackwater Brook. New Forest National Park. November 2014
Corncrake (Crex Crex) calling, with breath vapor in early morning, Kiskunsagi National Park, Hungary
Black-backed jackals (Canis mesomelas) two fighting with Plains / Burchell zebra (Equus burchelli) in foreground, Kalahari Desert, Nxai Pans National Park. Botswana
Spider webs covered in dew, on a flowering Gorse bush, Peak District National Park, UK October
Giant honey bee nest (Apis dorsata) up in a giant Mengaris tree (Koompassia excelsa) dipterocarp forest in Danum Valley, Sabah, Borneo, Malaysia
Kaieteur Falls is the worlds widest single drop waterfall, located on the Potaro river in the Kaieteur National Park, in Essequibo, Guyana, South America
Mud-covered face of bull African buffalo (Syncerus caffer) with a cateract in one eye, on the banks of the Chobe River, Chobe National Park, Botswana