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Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
Tufted / Brown capuchin (Cebus apella), female with baby on back, sitting in tree, mid-altitude montane forest, Manu Biosphere Reserve, Peru
Wild boar (Sus scrofa) eyes glowing at dusk, introduced species in La Pampa, Argentina
Warthogs (Phacochoerus aethiopicus) mother with babies drinking, Mkhuze game reserve, KwaZuiu-Natal, South Africa, June
Grykes in limestone pavement at sunset, , Twisteleton Scar End, Yorkshire Dales National Park, Yorkshire, England, UK, May 2013
European roller (Coracias garrulus) peering out of nest with feather, Lake Csaj, Pusztaszer, Kiskunsagi National Park, Hungary, February
African wild dog (Lycaon pictus) portrait, Mana Pools National Park, Zimbabwe
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
Mountain hare (Lepus timidus) in extreme winds with spin drifts of snow, Cairngorms National Park, Scotland, UK, February 2018
Scarletina bolete (Boletus luridiformis) toadstool, New Forest National Park, Hampshire, England, UK. September
Springbok (Antidorcas marsupialis) herd fleeing predator, Kgalagadi Transfrontier Park, South Africa
RF - Meerkat (Suricata suricatta) resting on cool sand, Kgalagadi Transfrontier Park, South Africa
Amazon kingfisher (Chloroceryle amazona) with fish, Cuiaba, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) with butterfly on snout, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) on river bank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male, hunting Capybara (Hydrochoerus hydrochaeris). The capybara while jumping away has cut the nose of the Jaguar with its toemail
Jaguar (Panthera onca) male, chasing a Caiman. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Lioness (Panthera leo) resting up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
Lion (Panthera leo) cub up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
African lion (Panthera leo) young male stalking towards camera, Etosha National Park, Namibia October
Female Bornean orangutan (Pongo pygmaeus) sitting in a tree, Tanjung Puting National Park, Borneo-Kalimantan, Indonesia
Black rhinoceros (Diceros bicornis) covered in mud, Mud on skin Etosha National Park, Namibia
Mountain gorilla (Gorilla beringei beringei) female resting with her baby, members of the Humba group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Mountain gorillas (Gorilla gorilla beringei) grooming, members of the Munyaga group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Moutain gorilla (Gorilla beringei beringei) close up of hand, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Portrait of male silverback Mountain gorilla (Gorilla beringei beringei) Virunga National Park, Democratic Republic of Congo
Mountain gorilla (Gorilla beringei) male silverback walking past, Susa group, Volcanoes National Park, Rwanda in wet season April
Mountain gorillas (Gorilla beringei) silverback with others, Agashya Group (Former 13 Group) Volcanoes National Park, Rwanda
Young Mountain gorilla (Gorilla beringei) this is Gakuru, one of 2 twin infants from female Kabatwa in Hirwa Group, on slopes of the Sabyinyo Volcano, Volcanoes National Park, Rwanda elevation 2610 m
Eastern lowland gorilla (Gorilla beringei graueri) silverback named Chimanuka, Kahuzi-Biega National Park, South Kivu Province, Democratic Republic of Congo (DRC)
Polar bear (Ursus maritimus) female coming out the den. Wapusk National Park, Churchill, Manitoba, Canada, March
Needle ice on Holly leaves (Ilex europaeus) in a woodland in Ambleside Cumbria UK. December 2008
Reindeer (Rangifer tarandus) bull in snow, Cairngorms Reindeer Herd, reintroduced to Scottish Highlands, Cairngorm National Park, Speyside, Scotland, December
Wild Rapa Nui horse / stallion, Rapa Nui National Park UNESCO World Heritage Site, Easter Island / Rapa Nui, Chile
Painted stork (Mycteria leucocephala), resting on nest with young at sunrise. Keoladeo / Bharatpur National Park UNESCO World Heritage Site, India
Jungle cat (Felis chaus), female in search of prey. Velavadar National Park, Gujarat, India
Asiatic wild buffalo (Bubalus arnee) portrait of female. Kaziranga National Park, India
Massive old growth oak trees of the cloud forest, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica. Small repro only
Mountain landscape of Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Alpine ibex (Capra ibex) close-up portrait. Gran Paradiso National Park, the Alps, Italy. January. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Crowned dwarf caiman (Paleosuchus trigonatus) male resting on a log. Yasuni National Park, Orellana, Ecuador
Land crab (Geosesarma sp. ) which raids Pitcher plant (Nepenthes ampullaria) for preyLand crab (Geosesarma sp.) which raids Pitcher plant (Nepenthes ampullaria) for prey, Sarawak, Borneo
Przewalski horse (Equus ferus przewalski) Khustain Nuruu National Park, Mongolia. December
Amur leopard (Panthera pardus orientalis) Land of the Leopard National Park, Primorsky Krai, Far East Russia. March
Chimpanzee (Pan troglodytes schweinfurthii) male, scratching its leg, National Park, Uganda
RF - Mountain gorilla (Gorilla beringei beringei) silverback male, portrait, member of the Nyakagezi group, Mgahinga National Park, Uganda. Critically endangered
Mountain gorilla (Gorilla beringei beringei) silverback male, portrait, member of the Nyakagezi group, Mgahinga National Park, Uganda. Critically endangered
Bluespine unicornfish (Naso unicornis) Shark Reef to Jolande, Ras Mohammed National Park, Egypt, Red Sea
