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Yacare caiman, (Caiman yacare) Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male on riverbank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) female, smelling scent marking of male. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) on river bank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male drinking, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
African lion cub (Panthera leo) in morning light. Moremi National Park, Okavango delta, Botswana
Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April
African lioness (Panthera leo) and cub, Etosha National Park, Namibia. January
RF- African Lion (Panthera leo) young male at sunrise, Etosha National Park, Namibia
Black rhino (Diceros bicornis) and dry grasses, Etosha National Park, Namibia
Black rhino (Diceros bicornis) approaching waterhole with her calf close by. Etosha National Park, Namibia
Angolan giraffes (Giraffa camelopardalis angolensis) and black rhinoceros (Diceros bicornis) reflecting in waterhole at sunset, Okaukuejo, Etosha National Park, Namibia, Africa
Black rhinoceros (Diceros bicornis) Etosha National Park, Namibia. March
RF- Black rhinoceros (Diceros bicornis) looking threatening, Etosha National Park, Namibia, June. Endangered species
Jaguar (Panthera onca) camera trap image, Tortuguero National Park, Costa Rica
Hoar frost on Beech leaves (Fagus sylvatica) Rydal, Lake District, England, UK. December 2010
Exmoor pony on Winsford Hill, Exmoor National Park, Somerset, England, UK. November 2013
Mountain range with rainforests in morning mist, Kudremukh National Park, Western Ghats UNESCO Natural World Heritage Site, India
Nilgai (Boselaphus tragocamelus) young bulls, Keoladeo / Bharatpur National Park UNESCO Natural World Heritage Site, India
Waterfall during monsoon, Kudremukh National Park, Western Ghats UNESCO Natural World Heritage Site, India. January 2016
Asiatic elephant (Elephas maximus), herd drinking water and crossing Mountain River at dawn, Jim Corbett National Park, India
Asiatic elephants (Elephas maximus), dust bathing at dawn. Jim Corbett National Park, India
Indian rhinoceros (Rhinoceros unicornis), female after light drizzle. Kaziranga National Park, India. March
Indian rhinoceros (Rhinoceros unicornis) in wetland, Kaziranga National Park, India
Black-crowned Central American squirrel monkey (Saimiri oerstedii) sitting on branch, Corcovado National Park, Osa Peninsula, Costa Rica, Vulnerable species
Puma (Puma concolor) resting, Corcovado National Park, Costa Rica, May
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Bairds tapir (Tapirus bairdii) walking along beach in Corcovado National Park, Costa Rica. Endangered
Leaf mantis (Choeradodis sp. ) Tortuguero National Park, Costa RicaLeaf mantis (Choeradodis sp.) Tortuguero National Park, Costa Rica
Giant owl butterfly, (Caligo memnon) Tortuguero National Park, Costa Rica
Eyespots on wing of Giant owl butterfly, (Caligo memnon) Tortuguero National Park, Costa Rica, February 2015
Ants (Formicidae) attacking newly emerged cicada (Cicadacae), Yasuni National Park, Ecuador. Highly commended in the Invertebrates Category of the Wildlife Photographer of the Year Awards (WPOY)
Guanaco (Lama guanicoe) with lake and mountains, Torres del Paine National Park, Patagonia. Chile
Cheetah male (Acinonyx jubatus) in morning light. Moremi National Park, Okavango delta, Botswana, Southern Africa
Underwater view of Sungai Melinau River, Gunung Mulu National Park, Sarawak, Borneo, Malaysia. Photographed for The Freshwater Project
Cheetah (Acinonyx jubatus) cub sitting portrait, Serengeti National Park, Tanzania
Rangimairewhenua or Blue Lake, sacred to the Maori, Nelson Lakes National Park, South Island, New Zealand, February 2013. Photographed for The Freshwater Project
Reindeer (Rangifer tarandus) bull reindeer with antlers in velvet, reintroduced Cairngorm Reindeer Herd, Cairngorm National Park, Speyside, Scotland, October
Lance-nosed chameleon (Calumma gallus), Andasibe-Mantadia National Park. Madagascar
RF - Indian rhinoceros (Rhinoceros unicornis) Kaziranga National Park, Assam, India
Hanuman langur (Semnopithecus / Presbytis entellus), female with baby at water tapHanuman langur (Semnopithecus/ Presbytis entellus), female with baby at water tap, Ranthambhore National Park, Rajasthan, India
Smooth-coated otter (Lutrogalle perspicillate) about to enter a marsh at dawn, Dudhwa National Park, Uttar Pradesh, India. Photographed using a camera trap
Fishing cat (Prionailurus viverrinus) walking along a path, Dudhwa National Park, Uttar Pradesh, India. Photographed using a camera trap
Kea (Nestor notabilis) juvenile in flight. Arthurs Pass National Park, South Island, New Zealand. Endangered Species
Ice washed up on Jokulsarlon, glacial lagoon, Skaftafell National Park, Iceland, February
Svartifoss waterfall, Skaftafell National Park, Iceland, February
Blood pheasant (Ithaginis cruentus) male, Baima Snow Mountain Nature Reserve, Yunnan, China, October
