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Red deer (Cervus elaphus) stag jumping over a stock fence along roadside, Cairngorms National Park, Scotland, UK. February
Atlantic salmon (Salmo salar) leaping up waterfall, Cairngorms National Park, Scotland, UK. October
Three female Putorana snow sheep (Ovis nivicola borealis) climbing up rocky mountainside with lamb following, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Female Grizzly bear (Ursus arctos horribilis) with cub among wildflowers, Grand Teton National Park, Wyoming, USA. June
Red fox (Vulpes vulpes) cub wandering through pine forest in golden glow of evening sunlight, Derbyshire, UK. August
Red Squirrel (Sciurus vulgaris) jumping over moss covered rock, NorthYorkshire, UK. June, 2021
Hundreds of Periodical cicada (Magicicada sp. ) nymphs ascending a tree trunk to metmorphose, Princeton, New JerseyHundreds of Periodical cicada (Magicicada sp.) nymphs ascending a tree trunk to metmorphose, Princeton, New Jersey, USA. June
Two Grey heron (Ardea cinerea) fighting over a fish, Pusztaszer reserve, Hungary. May
Purple heron (Ardea purpurea) walking along waters edge in early morning light, Pusztaszer reserve, Hungary. May
Eurasian otter (Lutra lutra) walking over partly frozen lake, Finland. March
Wolverine (Gulo gulo) walking through snow covered clearing, Finland. May
Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic, volumetric lighting, Hungary, January
Female Abyssinian ground hornbill (Bucorvus Abyssinicus) walking along riverbank, Allahein River, The Gambia
Western reef egret (Egretta gularis) running through shallow water with wings open, chasing small fish, Kotu creek, The Gambia
Stoat (Mustela erminea) on path, portrait, Strumpshaw Fen, Norfolk, UK. September
Coastal grey wolf (Canis lupus), British Columbia, Canada. August
Freshwater green hydra (Hydra viridissima) moving over a petri dish. Green colour comes from the symbiotic relationship with green alga Chlorella vulgaris
Weasel (Mustela erminea) running through deep snow with only its tail visible, Upper Bavaria, Germany, Europe. February
Dominant male Chimpanzee (Pan troglodytes troglodytes) walking in the mangrove, with female carrying her infant, Conkouati-Douli National Park, Republic of Congo, Africa
Juvenile Mute swan (Cygnus olor) in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Three Grey partridge (Perdix perdix) walking in snow. Near Nijmegen, the Netherlands. February
Humpback whale (Megaptera novaeangliae) diving into the depths, Hawaii, Pacific Ocean
Wild Wolverine (Gulo gulo) walking in snow, Mainua Kajaani, Finland. March
Sierra Negra giant tortoise (Chelonoidis guntheri) walking, Isabela Island, Galapagos, South America
Venomous Yellow-lipped sea krait (Laticauda colubrina) heading up to the surface to breathe, Philippines
Female Snow leopard (Panthera uncia) with her three month cub, BioParc Doue la Fontaine, France. Captive
Eurasian lynx (Lynx lynx) walking, Castilla La Mancha, Spain. August
Crescent-tail bigeye (Priacanthus hamrur) school on a coral reef, shot with a long exposure. North Male Atoll, Maldives. Indian Ocean
Common dolphin (Delphinus delphis) porpoising at the surface to breathe, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Male Jaguar (Panthera onca) walking through rainforest, Belize, Central America, 2017. Captive. Filmed for the BBC series Big Cats
Grizzly bear (Ursus arctos horribilis) running along shore of Yellowstone Lake at sunrise, Yellowstone National Park, Wyoming, USA
Two Yellow-eyed penguins (Megadyptes antipodes) emerging from the grasses, Enderby Island, Auckland Islands archipelago, New Zealand
Red squirrel (Sciurus vulgaris) leaping, Hawes, Yorkshire, England, UK, December
Proboscis monkey (Nasalis larvatus) male leaps between trees. Sabah, Malaysian Borneo
Comb-crested jacana (Irediparra gallinacea), Marrakai, Northern Territory, Australia, September
Hooded plover (Thinornis rubricollis) walking on beach looking for food. Skenes Creek, Victoria, Australia
Deepsea cirrate octopod {Stauroteuthis syrtensis} 830m, Gulf of Maine, Atlantic
Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
Roe deer (Capreolus capreolus) doe, leaping whilst running away, rear view, Vosges, France
A herd of wild konik horses running, with refection in water, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
RF - Red fox (Vulpes vulpes) walking through deep winter snow. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
Mallard (Anas platyrhynchos) shaking its wings with slow shutter speed to reveal movement, Martin Mere WWT Reserve, Lancashire, UK
RF - Dulmen pony, wild mares and foals running together at roundup. Dulmen, North Rhine-Westphalia, Germany. (This image may be licensed either as rights managed or royalty free.)
Arctic fox (Vulpes lagopus) walking in heavy snow. Hornstrandir Nature Reserve, Iceland. February
Roe deer (Capreolus capreolus) buck running past a garden shed at night, Wiltshire, UK, April. Property released
Flightless cormorant (Phalacrocorax harrisi) coming ashore amongst crabs. Clearwater Bay, Isabela Island, Galapagos. Sequence 3 of 3
Horses running in grassland. Bashang Grassland, near Zhangjiakou, Hebei Province, Inner Mongolia, China. July
RF-Horses running through Bashang Grassland. Near Zhangjiakou, Hebei Province, Inner Mongolia, China. July. (This image may be licensed either as rights managed or royalty free)
