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Leopard (Panthera pardus) female, walking across grassland in morning sunlight, Kenya
Eastern rockhopper penguin (Eudyptes chrysocome filholi) mid-fall after jumping off rocky ledge to get to sea, surrounded by Bull kelp (Durvillaea antarctica), Auckland Island, Auckland Island
Brown bear (Ursus arctos) female with two cubs walking through misty, woodland meadow, Finland. June
Arctic fox (Alopex lagopus) in winter coat, walking across snow, Svalbard, Norway. April
Polar bear (Ursus maritimus) female with cub, walking over snow with mountains in background, Svalbard, Norway. April
Giant anteater (Myrmecophaga tridactyla) walking, Pantanal, Mato Grosso do Sul, Brazil
Iberian ibex (Capra pyrenaica) kid, aged two months, making its way down cliff, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), adult female, walking along coastal cliff. Mediterranean coast, Andalusia, Spain. November
African elephant (Loxodonta africana), female and calf, crossing plain at sunset. Amboseli National Park, Kenya. July
Masai giraffe (Giraffa camelopardalis tippelskirchi) running across plain. Amboseli National Park, Kenya. July. Digitally enhanced
Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp)Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp.) being followed by a California sea lion (Zalophus californianus), Espiritu Santo Island
Water rail (Rallus aquaticus) wading through water, Dumfries and Galloway, Scotland, UK
Sand cat (Felis margarita) walking over sandy ground. Captive, occurs in North Africa, the Arabian Peninsula, Pakistan and the Middle East
Female Olive ridley turtles (Lepidochelys olivacea) coming ashore to nest while others leaves, at sunrise, during massive arribada. Playa Escobilla Sanctuary, Oaxaca, Mexico. Pacific Ocean
Adult American oystercatcher (Haematopus palliatus) walking along beach, with chick running and flapping its wings. Long Island, New York, USA. June
Gemsbok (Oryx gazella) female and calf walking past sand dune, Sossusvlei UNESCO World Heritage Site, Namib desert, Namib-Naukluft National Park, Namibia
Herd of wild Onaqui horses (Equus ferus) trotting in dust. Onaqui Mountain Herd Management Area, Utah, USA. September
Brown bear (Ursus arctos) walking across rock in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. September
Sika deer (Cervus nippon) stag walking on mountain slope at the edge of woodland, with forested mountains in the background, Land of the Leopard National Park, Russian Far East
Sika deer (Cervus nippon) herd walking past fallen tree in forest as one looks around, Land of the Leopard National Park, Russian Far East. Taken with remote camera. June
Amur leopard (Panthera pardus orientalis) walking past snow-covered trees on mountain slope, Land of the Leopard National Park, Russian Far East. Critically endangered. Taken with remote camera
Sika deer (Cervus nippon) doe walking across snow covered mountain slope, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Chocolate Labrador retriever puppy running on garden lawn, Rhode Island, USA. May
Puma (Puma concolor) female, walking in deep snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Puma (Puma concolor) cub, aged nine months, chasing scavenging White-throated Caracaras (Phalcoboenus albogularis), Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Three Guanacos (Lama guanicoe) running over hillside in front of the Towers granite rock formation and low cloud, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female, running sequence in deep snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile. Composite of five separate images
Two Sockeye / Red Salmon (Oncorhynchus nerka), female digging riverbed to lay eggs on spawning ground. Trees showing autumnal colours, Adams river, British Columbia, Canada. October
Two Leatherback turtle (Dermochelys coriacea) hatchlings moving across beach towards sea after leaving nest, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean Sea
Hanuman langur (Presbytis entellus) juvenile, running up tree trunk, Bandhavgarh National Park, India
Cockapoo puppy, aged 18 weeks, running, portrait
Cockapoo puppy, aged 4 months, running, portrait
Norfolk terrier puppy, aged 7 months, running, portrait
Galapagos sea lion (Zalophus wollebaeki) juvenile, jumping and playing in ocean, San Cristobal Island, Galapagos National Park, Pacific Ocean. Endangered
Puma (Puma concolor) cub running, Torres del Paine National Park, Magallanes, Chile
Herd of African elephants (Loxodonta africana) coming to drink at waterhole. Tsavo East National Park, Kenya
Wolf (Canis lupus) walking in forest, Finland. September
Spinner dolphin (Stenella longirostris) juvenile surfacing next to its mother, Sri Lanka
Indri (Indri indri) male, leaping through the rain forest canopy, Andasibe-Mantadia National Park, eastern Madagascar. Critically Endangered
Wood duck (Aix sponsa) female, leading brood of ducklings on pond, Acadia National Park, Maine, USA. June
Black grouse (Lyrurus tetrix) male, jumping, part of a dominance display, at lek in early morning light, Kuusamo, Finland, April
Arctic fox (Vulpes lagopus) hunting along rocky shoreline under overhanging ice, Svalbard, Norway. July
Brown hare, (Lepus europaeus), male and female displaying courting behaviour, Islay, Scotland, UK. March
Brown hare, (Lepus europaeus), males in pursuit of female that is in season, Islay, Scotland, UK. March
Crested caracara (Caracara cheriway), adult, walking on beach. Pacific coast, Oaxaca state, Mexico. November
Group of Clarion angelfish (Holacanthus clarionensis) cleaning an Oceanic manta ray (Mobula birostris) as it swims over the reef, Revillagigedo Islands, Mexico, Pacific Ocean
Greater one-horned rhinoceros, (Rhinoceros unicornis) walking through shallow water, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis), small herd leaping through grass, Bardia National Park, Terai, Nepal