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Male Yellow boxfish (Ostracion cubicus) displaying, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Female Jaguar (Panthera onca) playing with her cub, captive, occurs in Southern and Central America
Lion (Panthera leo) male smelling, flehmen response, Masai-Mara Game Reserve, Kenya. Vulnerable species
Red Foxes (Vulpes vulpes) young play fighting. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Wilsons Storm Petrel (Oceanites oceanicus) feeding whilst walking on water Atlantic ocean, Madeira Portugal, August
Whale shark (Rhincodon typus) mouth open filter feeding at the surface, Quintana Roo, Yucatan Peninsular, Mexico, Caribbean Sea
Yacare Caiman (Caiman yacare) with mouth open to keep cool, Pantanal, Brazil
Lemon sole (Microstomus kitt) portrait. Gardur, south west Iceland. North Atlantic Ocean
Red deer (Cervus elaphus) stag bellowing. Surrey, UK, October
Robin (Erithacus rubecula) singing, Norfolk, England, UK, February
RF- Wide angle portrait of Hairy frogfish (Antennarius striatus) waiting to ambush prey on black sand slope. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Portrait of Male Sockeye salmon (Oncorhynchus nerka) showing teeth. Adams River, British Columbia, Canada, October
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Spectacled caiman (Caiman crocodilus) Rio BaiAia Bonita, Bonito, Mato Grosso do SulSpectacled caiman (Caiman crocodilus) Rio Baiia Bonita, Bonito, Mato Grosso do Sul, Brazil
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills, Lake Kerkini, Greece, February. Highly Commended, 2011 Wildlife Photographer of the Year competition
RF- Male Bengal Tiger (Panthera tigris tigris) face portrait in undergrowth, Kanha National Park, Madhya Pradesh, India. Endangered species
Close up of Konik wild horse (Equus ferus caballus) snout, Meinerswijk nature reserve, near Arnhem, the Netherlands. May
Eastern Hermann's tortoise (Testudo hermanni boettgeri) hatching from egg, mouth open. Captive, occurs in South East Europe
Male Lemon shark (Negaprion brevirostris) gliding just below the surface, Grand Bahama, Caribbean Sea
Common dab (Limanda limanda) portrait, Trondheimfjord, Norway, North Atlantic Ocean
Robin (Erithacus rubecula) male singing, perched on branch. Joutsa, Finland. April
Cunner (Tautogolabrus adspersus) pair in Eelgrass (Zostera marina) bed. Newfoundland, Canada. September
Ring-tailed lemur (Lemur catta) suckling baby. Berenty Reserve, Madagascar
Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
Black-bellied bustard (Lissotis melanogaster) male calling at sunrise. Serengeti National Park, Tanzania
African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia
Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) on sea, tossing fish. Turtle Cove, Santa Cruz Island, Galapagos
Raccoon (Procyon lotor) approaching with curiousity. Stanley Park, Vancouver, British Columbia, Canada. August
Silky sifaka (Propithecus candidus) female with baby sitting amongst branches in understorey of mid-altitude montane rainforest. Marojejy National Park, north east Madagascar
RF- Stump tailed chameleon (Brookesia superciliaris) portrait, Andasibe-Mantadia National Park, Eastern Madagascar
Canary fish (Epinephelus marginatus) in reef. El Hierro, Canary Islands
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Hawfinch 1+Coccothraustes coccothraustes+2 perched on branch amongst Hawthorn 1+Crataegus sp+2 berries. Sierra de Grazalema Natural Park, Cadiz, Spain. February
Bengal tiger (Panthera tigris), portrait. Bandhavgarh National Park, India. Photo© Phillip Ross/Felis Images
Artichoke coral (Scolymia cubensis), close-up of mouth. Gardens of the Queen National Park, Cuba
Ornamental koi carp (Cyprinus carpio) group with open mouths at lake surface. Xishuangbanna Tropical and Flowers Garden, Jinghong, Yunnan, China
Gannet (Morus bassanus) chick with fluffy with mouth open. Shetland Islands, Scotland, UK, August
American crocodile (Crocodylus acutus) with mouth open over seagrass bed, Chinchorro Banks Biosphere Reserve, Quintana Roo, Mexico
Cormorants (Phalacrocorax carbo) squabbling over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Whale shark (Rhincodon typus) front view portrait, Cenderawasih Bay, West Papua. Indonesia
Grey seal (Halichoerus grypus) young male plays and gapes, showing his teeth. Lundy IslandGrey seal (Halichoerus grypus) young male plays and gapes, showing his teeth.Lundy Island, Devon, UK, Bristol Channel, August
Golden eagle (Aquila chrysaetos) male head portrait beak open calling, Kalvtrask, Vasterbotten, Sweden. December
Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
