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Large male Napoleon wrasse (Cheilinus undulatus) yawning. Ras Mohammed, Sinai, Egypt. Red Sea
Male Yellow-headed jawfish (Opistognathus aurifrons) blows out the clutch of eggs he is incubating in his mouth to oxygenate them. East End, Grand Cayman, Cayman Islands, British West Indies
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RF - California sea lion (Zalophus californianus) with mouth open close to the camera. The milk teeth of this pup show decay, which is quite usual. Los Islotes, La Paz, Baja California Sur, Mexico
American crocodile (Crocodylus acutus), underwater, Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January. Vulnerable species
Frilled shark (Chlamydoselachus anguineus) specimen with mouth open, from Atlantic Ocean, at a depth of 729m
Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil
Black howler monkeys (Alouatta caraya) male and two females calling from tree, captive, Apenheul Park, Netherlands
Loggerhead sea turtle (Caretta caretta) emerging from shell, Dalyan delta, Turkey, July
Humpback whale (Megaptera novaeangliae) portrait, Moorea, French Polynesia, Pacific Ocean
Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Blotched blue-tongue skink (Tiliqua nigrolutea) female gaping with tongue out, in defense against perceived threat. Lake Tantangara, Snowy Mountains, New South Wales, Australia
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
Peregrine falcon (Falco peregrinus) in flight with open beak. The Netherlands. May
Tiger moray (Enchelycore anatina) portrait, Tenerife, Canary Islands
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Fangtooth moray (Enchelycore anatina), close up of eye and open mouth with teeth. Tenerife, Canary Islands
Galapagos sea lion (Zalophus wollebaeki) jumping out of Pacific ocean. Sombrero Chino, Santiago Island, Galapagos
Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December
American crocodile (Crocodylus acutus) with open mouth amongst Seagrass (Alismatales). Gardens of the Queen National Park, Cuba
Giant panda (Ailuropoda melanoleuca) dusted with snow eating Bamboo. In natural enclosure, Panda Breeding Centre, Wolong, Sichuan Province, China. February 2008
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
Galapagos land iguana (Conolophus subcristatus) portrait with mouth open, Galapagos
Emu (Dromaius novaehollandiae) juvenile with beak open to catch a leaf, Cleland Wildlife Park, South Australia, March. Captive
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May
Reef manta (Manta alfredi) mouth open feeding at night, South Ari atoll, Maldives, Indian Ocean
Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Jaguar (Panthera onca) female snarling, native to Southern and Central America, captive
Eastern green mambas (Dendroaspis angusticeps) one with mouth wide open, captive, from East Africa
Sperm whale (Physeter macrocephalus) with mouth open, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean
Black skimmer (Rynchops niger) foraging along surf line. Tierra Verde, Florida, USA
Hippopotamus (Hippopotamus amphibius) in pool with mouth open. Mana Pools National Park, Zimbabwe
Antarctic fur seal (Arctocephalus gazella), mother warning off curious juvenile. Southern elephant seal (Mirounga leonina) in background. Gold Harbour, South Georgia. October
Lionesses (Panthera leo) grooming each other, Masai-Mara Game Reserve, Kenya
Arctic fox (Vulpes lagopus) in summer coat calling, Wrangel Island, Far Eastern Russia, August
Tokay gecko (Gekko gecko) Pat Sin Leng Country Park, New Territory, Hong Kong, China
Fawn Pug dog and puppy, 8 weeks
Greylag goose (Anser anser) taking flight from lake. Southern Norway. March
Great white shark (Carcharodon carcharias) Guadalupe Island or Isla Guadalupe, Pacific Ocean, Mexico, September. Vulnerable species
Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
Grasshopper Warbler (Locustella naevia) singing, Joensuu, Finland, May
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Male Yellow boxfish (Ostracion cubicus) displaying, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Female Jaguar (Panthera onca) playing with her cub, captive, occurs in Southern and Central America