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Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean
Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
Portrait of a Golden snub-nosed monkey (Rhinopithecus roxellana) screaming and showing its teeth, Foping Nature Reserve, Shaanxi, China. Endangered species
Close up portrait of the face of a Nurse shark (Ginglymostoma cirratum) resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas
Trap-jaw ant (Odontomachus hastatus) with mandibles open, Los Amigos Biological Station, Peru
Grey whale (Eschrichtius robustus) at water surface with mouth open showing baleen plates, San Ignacio Lagoon, Baja California, Mexico
Arctic Fox (Alopex / Vulpes lagopus) yawning, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Tree sparrow (Passer montanus) feeding a fledgling on a branch of Hawthorn (Crataegus monogyna) blossom, Perthshire, Scotland. June 2013
RF - Nile crocodile (Crocodylus niloticus head close up with jaws open)RF - Nile crocodile (Crocodylus niloticus head close up with jaws open, Chobe river, Botswana. (This image may be licensed either as rights managed or royalty free.)
Guineafowl puffer fish (Arothron meleagris) inflated, displaying defensive behaviour over coral reef, Yap, Indo Pacific, Micronesia
Salmon shark (Lamna ditropis). Port Fidalgo, Alaska, USA. July
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
Keel-billed toucan (Ramphastos sulfuratus) feeding, tossing fruit seed in beak, perched on branch. Boca Tapada, Costa Rica. Sequence 2/2
Walrus (Odobenus rosmarus) amongst group hauled out on shore. Spitsbergen, Svalbard, Norway. July
Tokay gecko (Gekko gecko) enacting a defensive display towards a perceived threat, Phuket Island, Thailand, March. Controlled conditions
RF-Asian elephant (Elephas maximus) dust bathing. Jim Corbett National Park, Uttarakhand, India. (This image may be licensed either as rights managed or royalty free)
Javan leopard (Panthera pardus melas) roaring, native to the Indonesian island of Java. Captive. Digital composite
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Smalltooth sand tiger shark (Odontaspis ferox), view from below. El Hierro. Canary Islands
Fangtooth moray (Enchelycore anatina) with open mouth, portrait. Tenerife, Canary Islands
Napoleon wrasse (Cheilinus undulatus) on a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October
Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December
Aerial view of Fin whale (Balaenoptera physalus) lunge-feeding, with mouth open and throat pouch distended, southern Sea of Cortez (Gulf of California), Baja California, Mexico
Emu (Dromaius novaehollandiae) juvenile with beak open to catch a leaf, Cleland Wildlife Park, South Australia, March. Captive
Yorkshire Terrier mother, Evie, with her two Yorkishire terrier x Poodle Yorkipoo pups, age 7 weeks
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Arctic fox (Vulpes lagopus), juvenile licking lips, winter pelage. Dovrefjell National Park, Norway. September
Nurse shark (Ginglymostoma cirratum) portrait, resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas. The Bahamas National Shark Sanctuary
Nile crocodile (Crocodylus niloticus) Chobe National Park, Botswana, June
Indonesian blue-tongued skink (Tiliqua gigas) with tongue extended, captive, from Papua New Guinea
Great white shark (Carcharodon carcharias) breaking surface with mouth open. Guadalupe Island, Mexico
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
Mouth of Common Sea Urchin (Echinus esculentus) Guillaumesse, Sark, British Channel Islands
Tasseled wobbegong shark (Eucrossorhinus dasypogon) yawning. Citrus Ridge, Yanggefo Island, Gam Island, Raja Ampat, West Papua, Indonesia
Turbot (Scophthalmus maximus) detail of the eyes and mouth, United Kingdom, September
Humpback whale (Megaptera novaeangliae) portrait, Moorea, French Polynesia, Pacific Ocean
Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Blotched blue-tongue skink (Tiliqua nigrolutea) female gaping with tongue out, in defense against perceived threat. Lake Tantangara, Snowy Mountains, New South Wales, Australia
Peregrine falcon (Falco peregrinus) in flight with open beak. The Netherlands. May
Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia
Tiger moray (Enchelycore anatina) portrait, Tenerife, Canary Islands
Raccoon (Procyon lotor) approaching with curiousity. Stanley Park, Vancouver, British Columbia, Canada. August
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Fangtooth moray (Enchelycore anatina), close up of eye and open mouth with teeth. Tenerife, Canary Islands
Galapagos sea lion (Zalophus wollebaeki) jumping out of Pacific ocean. Sombrero Chino, Santiago Island, Galapagos
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Artichoke coral (Scolymia cubensis), close-up of mouth. Gardens of the Queen National Park, Cuba
American crocodile (Crocodylus acutus) with open mouth amongst Seagrass (Alismatales). Gardens of the Queen National Park, Cuba
Ornamental koi carp (Cyprinus carpio) group with open mouths at lake surface. Xishuangbanna Tropical and Flowers Garden, Jinghong, Yunnan, China
Giant panda (Ailuropoda melanoleuca) dusted with snow eating Bamboo. In natural enclosure, Panda Breeding Centre, Wolong, Sichuan Province, China. February 2008
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
Galapagos land iguana (Conolophus subcristatus) portrait with mouth open, Galapagos
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May
American crocodile (Crocodylus acutus) with mouth open over seagrass bed, Chinchorro Banks Biosphere Reserve, Quintana Roo, Mexico
Whale shark (Rhincodon typus) front view portrait, Cenderawasih Bay, West Papua. Indonesia
Reef manta (Manta alfredi) mouth open feeding at night, South Ari atoll, Maldives, Indian Ocean
Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Jaguar (Panthera onca) female snarling, native to Southern and Central America, captive
Eastern green mambas (Dendroaspis angusticeps) one with mouth wide open, captive, from East Africa
Sperm whale (Physeter macrocephalus) with mouth open, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
Black skimmer (Rynchops niger) foraging along surf line. Tierra Verde, Florida, USA
Hippopotamus (Hippopotamus amphibius) in pool with mouth open. Mana Pools National Park, Zimbabwe
Close up of Konik wild horse (Equus ferus caballus) snout, Meinerswijk nature reserve, near Arnhem, the Netherlands. May
Eastern Hermann's tortoise (Testudo hermanni boettgeri) hatching from egg, mouth open. Captive, occurs in South East Europe
Male Lemon shark (Negaprion brevirostris) gliding just below the surface, Grand Bahama, Caribbean Sea
Common dab (Limanda limanda) portrait, Trondheimfjord, Norway, North Atlantic Ocean
Robin (Erithacus rubecula) male singing, perched on branch. Joutsa, Finland. April
Cunner (Tautogolabrus adspersus) pair in Eelgrass (Zostera marina) bed. Newfoundland, Canada. September
Ring-tailed lemur (Lemur catta) suckling baby. Berenty Reserve, Madagascar
Black-bellied bustard (Lissotis melanogaster) male calling at sunrise. Serengeti National Park, Tanzania
Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) on sea, tossing fish. Turtle Cove, Santa Cruz Island, Galapagos
Silky sifaka (Propithecus candidus) female with baby sitting amongst branches in understorey of mid-altitude montane rainforest. Marojejy National Park, north east Madagascar
RF- Stump tailed chameleon (Brookesia superciliaris) portrait, Andasibe-Mantadia National Park, Eastern Madagascar
Canary fish (Epinephelus marginatus) in reef. El Hierro, Canary Islands
Hawfinch 1+Coccothraustes coccothraustes+2 perched on branch amongst Hawthorn 1+Crataegus sp+2 berries. Sierra de Grazalema Natural Park, Cadiz, Spain. February
