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Sohal surgeonfish (Acanthurus sohal), Red Sea, Egypt. January
Southern Sea otter (Enhydra lutris) mother holding young pup (less than one week in age), Monterey Bay, California, USA
Sea otter (Enhydra lutris) floating on its back at the surface among the kelp, Alaska, USA, Gulf of Alaska. Pacific ocean
Sea otter (Enhydra lutris) grooms itself in icy waters off Columbia Glacier in Prince William Sound, Chugach National Forest, Alaska, USA, May
Southern Elephant Seals (Mirounga leonina) pup with mother. King Haakon Bay, South Georgia. November
Southern Elephant Seal bull (Mirounga leonina) close up portrait, King Haakon Bay, South Georgia. November
Coral formations at low tide in Prony Bay in the Southern Lagoon, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Griffon vulture (Gyps fulvus) landing on coast next to flock. Cantabria, Spain, March
Galapagos green turtle (Chelonia mydas agassizi) swimming in coastal waters of San Cristobal Island, Galapagos
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Sea otter (Enhydra lutris) resting amongst sea ice, Alaska, USA, June
Sperm whale (Physeter macrocephalus) aerial view. Baja California, Mexico
Short-finned pilot whale (Globicephala macrorhynchus) aerial view, Baja California, Mexico
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Lemon shark pups (Negaprion brevirostris) spend the first 5-8 years of their life in mangrove forests. The tangle of roots provides protection from predators
Part of a submerged Holocene forest, revealed after a winter storm from beneath the sand at Hells Mouth (Porth Neigwl), Abersoch Wales, UK March 2014
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Hard corals (Acropora microclados) growing in shallow water, reflected in the surface, Gubal Island, Egypt. Strait of Gubal, Red Sea
Munks mobula ray / Devilray (Mobula munkiana) leaping out of the water, Sea of Cortez, Gulf of California, Baja California, Mexico, February
Bohar snapper (Lutjanus bohar) school at Ras Mohammed, Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
Whale Shark (Rhincodon typus) and Golden trevally (Gnathanodon speciosus) Cenderawasih Bay, West Papua, Indonesia
Killer whale (Orcinus orca) hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Reef manta (Manta alfredi) mouth open feeding at night, South Ari atoll, Maldives, Indian Ocean
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Historical illustration of Common octopus (Octopus vulgaris) Pierre Denys de Montfort engraving from Histoire Naturelle Generale et Particuliere des Mollusques, 1801-1802
Yellow warty frogfish (Antennarius maculatus) Seraya, Tulamben area, Bali, Indonesia. Java Sea
Aerial view of small coral island, Western Provinces, Solomon Islands
Common eider (Somateria mollissima), female on her nest with sea behind, Finland, May
Southern Rockhopper penguin (Eudyptes chrysocome) colony, Kidney Island, Falkland Islands, October. Vulnerable species
Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March
Common dolphin (Delphinus delphis) close up of dolphin surfing at speed, Hebrides, Scotland, UK
Sea cucumber / Sea apple (Pseudocolochirus violaceus) Rinca, Komodo National Park, Indonesia
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Galapagos Shark (Charcharhinus galapagensis), Roca Partida islet, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Striped Dolphin (Stenella coeruleoalba) in the strait of Gibraltar, Spain
Southern elephant seal (Mirounga leonina), fight between two males, Saint Andrew, South Georgia
Nurse sharks (Ginglymostoma cirratum) over seagrass at night. White spiracle behind eye is visible. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Sea apple (Pseudocolochirus violaceus) Komodo National Park, Indonesia
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean
Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
School of Munks devil ray (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Sea otter (Enhydra lutris) mother and sleeping newborn pup (aged 3 days) Monterey, California, USA. *Digitally darkened scar on mothers nose
Fluorescent coral photographed with ultraviolet / UV light, Bonaire, Leeward Antilles, Caribbean region, Netherlands Antilles
Patagonian redfish (Sebastes oculatus) about 4cm, Comau Fjord, Patagonia, Chile
Common lobster (Homarus gammarus) looks out from its hole amongst Dead mans fingers (Alcyonium digitatum) beneath a forest of Oarweed kelp (Laminaria hyperborea)
