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American crocodile (Crocodylus acutus) with open mouth amongst Seagrass (Alismatales). Gardens of the Queen National Park, Cuba
Orca (Orcinus orca) pod hunting Sealion, tossing it out of the sea, Punta Norte, Valdez Peninsula, Argentina. April
Orca (Orcinus orca) hunting South American sealion (Otaria flavescens) close to shore. Punta Norte, Valdez Peninsula, Argentina. April
Shoal of colorful yellow fish Panamic porkfish (Anisotremus taeniatus), Cabo Pulmo Marine National Park, Baja California Sur, Mexico
Shoal of Dog snapper, (Lutjanus novemfasciatus) in front of shoal of Big-eye jacks (Caranx sexfasciatus), Cabo Pulmo Marine National Park, Baja California Sur, Mexico
Little penguin (Eudyptula minor) standing on rocks of St Kilda breakwater, returning to nest at dusk. Melbourne city in background. Victoria, Australia. December 2016
Wandering albatross (Diomedea exulans) flying on the open ocean, Drake passage, Antarctic Peninsula, Antarctica
Bristle worm (Polychaeta sp), Antarctic Peninsula, Antarctica
Sea angel (Clione limacina), Antarctic Peninsula, Antarctica
Leopard seal (Hydrurga leptonyx), Antarctic Peninsula, Antarctica
Weddell seal (Leptonychotes weddellii) hauled out on ice, Antarctic Peninsula, Antarctica
Halimeda ghost pipefish (Solenostomus halimeda) mimicking the halimeda algae. Sulu sea, Philippines
False clown anemonefish (Amphiprion ocellaris) in a magnificent sea anemone (Heteractis magnifica), Sulu sea, Philippines
Coral reef scene filled with a great diversity of fish and coral life, near Alor, Indonesia
RF - Reef manta (Mobula alfredi) female swimming close to a coral reef, while Cleaner wrasse (Labroides dimidiatus), tiny by comparison, pick parasites from its belly
RF - Silhouette of a California sea lion (Zalophus californianus) swimming through an underwater cave. Los Islotes, La Paz, Baja California Sur, Mexico
RF - Portrait of Coral hermit crab (Paguritta harmsi) living in a hard coral colony. North Male Atoll, Maldives. Indian Ocean (This image may be licensed either as rights managed or royalty free.)
RF - A school of Crescent-tail bigeye (Priacanthus hamrur) on a coral reef. North Male Atoll, Maldives. Indian Ocean (This image may be licensed either as rights managed or royalty free.)
RF - Basking shark (Ceterhinus maximus) feeding on plankton in Senen Cove, Cornwall, UK. June. (This image may be licensed either as rights managed or royalty free.)
RF - Denise's pygmy seahorse (Hippocampus denise) looks out from its home in a sea fan (Annella sp.). Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Portrait of a Bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Scalloped hammerhead shark (Sphyrna lewini) swims through a school of Pacific creolefish (Paranthias colonus). Wolf Island, Galapagos National Park, Galapagos Islands.. East Pacific Ocean
Portrait of a tarpon (Megalops atlanticus) on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Portrait of a male short snouted seahorse (Hippocampus hippocampus) in sea oak seaweed (Halidrys siliquosa). Devon, England, United Kingdom. British Isles. English Channel. North East Atlantic Ocean
Coral (Lobophytum sp. ) on a reef flat, with Hawksbill turtle (Eretmochelys imbricata)Coral (Lobophytum sp.) on a reef flat, with Hawksbill turtle (Eretmochelys imbricata) in background. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Blacktip reef shark (Carcharhinus melanopterus) swims through schools of fish. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Killer whales (Orcinus orca) at the surface in an Arctic fjord in Spildra, northern Norway, with sea birds. Arctic Ocean
California sea lion (Zalophus californianus) pups in an underwater cave. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean
Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind, in the plankton-rich green waters of La Paz bay
Tawny nurse sharks (Nebris ferrungineus) at night. Vavuu Atoll, Maldives. Indian Ocean
Several mimic goatfish (Mulliodichthys mimicus) hide within a school of Bluestripe snapper (Lutjanus kasmira) on a coral reef. North Ari Atoll, Maldives. Indian Ocean
School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean
Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the packground sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia
Thorny seahorse (Hippocampus histrix) gripping onto a sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Veined octopus (Amphioctopus marginatus) emerging from its home in an old clam shell. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Sleeping sperm whale, (Physeter macrocephalus) Dominica, Caribbean Sea, Atlantic Ocean
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Marine iguana (Amblyrhynchus cristatus), Punta Vicente Roca, Isabela Island, Galapagos
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Galapagos sea lions (Zalophus wollebaeki) fighting, Galapagos
Golden cownose rays (Rhinoptera steindachnyri) schooling in sheltered mangrove lagoon, Academy Bay, Santa Cruz Island, Galapagos, Ecuador
Long-finned pilot whale (Globicephala melas) off Nova Scotia, Canada. October
Atlantic bluefin tuna (Thunnus thynnus) off Antigonish, Nova Scotia, Canada. October
Overhead view of a blue shark (Prionace glauca) off Halifax, Nova Scotia, Canada. July
Ocean sunfish (Mola mola) off Halifax, Nova Scotia, Canada. July