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Hydromedusa / Trachymedusa (Crossota sp) from approx 2700m, mid Atlantic ridge, June 2010Hydromedusa/ Trachymedusa (Crossota sp) from approx 2700m, mid Atlantic ridge, June 2010
Deepsea octopus (Grimpoteuthis sp), Dumbo, North Atlantic
White-tailed sea eagle (Haliaeetus albicilla) flying over water with male Common eider prey (Somateria mollissima), Uto, Parainen, Finland, April
Group of Bigfin reef squids (Sepioteuthis lessoniana) swimming at night, Indonesia, Sea of Flores
Bigfin reef squid (Sepioteuthis lessoniana) swimming at night, Indonesia, Sea of Flores
Brown fur seal / Cape fur seal (Arctocephalus pusillus), Western Cape, South Africa. Atlantic Ocean
Palmers globemallow (Sphaeralcea palmeri) and view towards Toro Islet and Guadalupe Island, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, September
Day octopus (Octopus cyanea) female, in mid-water, Hawaii
Red mangrove (Rhizophora mangle) propagules / plantlets which become fully mature plants before dropping off the parent tree to drift away and establish a new tree. Bahamas
Queen conch (Aliger gigas) in a seagrass (Thalassia testudinum) meadow, Bahamas
Red sea fan (Paramuricea clavata) with gorgonian corals and Light-bulb sea squirt (Clavelina lepadiformis) Punta Carena, Capri Island, Sorrentine Peninsula, Italy, Tyrrhenian Sea, Mediterranean
Crinoid or feather star (Antedon mediterranea) on Violescent sea whip or Red sea fan (Paramuricea clavata) Punta Campanella Marine Protected Area, Amalfi Coast, Italy, Tyrrhenian Sea, Mediterranean
Slate pencil sea urchins (Heterocentrotus mammillatus) on coral reef with Pyramid butterflyfish (Hemitaurichthys polylepis), Hawaii
Sea anemone (Calliactis parasitica) usually associated with hermit crabs, here on a Mediterranean bonnet shell (Semicassis granulata), Ponza island, Italy, Tyrrhenian Sea, Mediterranean
Loggerhead turtle (Caretta caretta) accompanied by pilotfish (Naucrates ductor) swimming near the surface. Santa Maria Island, Azores, Portugal, Atlantic Ocean. Vulnerable
Low tide on Westward Ho! beach, tinged pink at sunrise. Devon, England, UK. February 2020
Thrift (Armeria maritima) flowering on cliff top at sunset. Hartland Quay, Devon, England, UK. May 2019
The Green Bridge of Wales sea arch and stack along limestone coastline. Castlemartin, Pembrokeshire, Wales, UK, September 2019
Atlantic spotted dolphins, (Stenella frontalis), Formigas Islet dive site, Santa Maria Island, Azores, Portugal, Atlantic Ocean
Spotted gully shark (Triakis megalopterus) in coral reef. Simons Town, Western Cape, South Africa. June
Scoophead shark (Sphyrna media), released specimen. Isla Chepillo, Rio Bayano, Panama
Pyjama shark (Poroderma africanum) in reef. Simons Town, Western Cape, South Africa. June
Pyjama shark (Poroderma africanum), two in reef, amongst kelp. Simons Town, Western Cape, South Africa. May
Japanese bullhead shark (Heterodontus japonicus) on reef. Chiba Prefecture, Honshu, Japan. May
Eastern shovelnose ray (Aptychotrema rostrata) on sea floor. Nelson Bay, New South Wales, Australia. July
Cuban dogfish (Squalus cubensis). Exuma Sound, Bahamas
Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Close up at the dental plates of Three color parrotfish (Scarus tricolor) as it comes in for a bite of hard coral, Fiji
Banded sea krait (Laticauda colubrina) hunting in seagrass bed. Manado, North Sulawesi, Indonesia
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
Neptune seagrass (Posidonia oceanica) bed, sun rays shining through water. A patch of seagrass bed in the Mediterranean sea is considered to be the oldest living organism on earth. Spain. June
Atlantic blue crab (Callinectes sapidus) hunting in Turtlegrass (Thalassia testudinum) seagrass bed, view towards sky. Florida Keys, Florida, USA
Orange and white soft corals, Dead mans fingers (Alcyonium digitatum) thriving beneath kelp (Laminaria hyperborea). St Abbs, Eyemouth, Berwickshire, Scotland, United Kingdom. North Sea
Cuttlefish (Sepia officinalis) male hovers over sea oak (Halidrys siliquosa). Babbacombe, Torquay, Devon, England, United Kingdom. English Channel. North East Atlantic Ocean
RF - Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and Sea plumes (Pseudopterogorgia sp)
Green sea turtle (Chelonia mydas) near surface in shallow water. The Bahamas
Caribbean reef squid (Sepioteuthis sepioidea) eating fish at night, portrait. The Bahamas
Spiny seahorse (Hippocampus guttulatus) adult female in a meadow of (Zostera marina) seagrass. Studland Bay, Dorset, England, United Kingdom. English Channel, North East Atlantic Ocean
White and yellow nudibranch (Polycera quadrilineata) feeding on a sea mat (Membranipora membranacea). Shetland Islands, Scotland, British Isles. United Kingdom. North East Atlantic Ocean
School of Oriental sweetlips (Plectorhinchus vittatus) gather tightly together as they rest during the day, with a Clown triggerfish (Balistoides conspicillum)
Black rockfish (Sebastes melanops) school shelter in a bull kelp forest (Nereocystis luetkeana). Race Rocks, Victoria, Vancouver Island, British Columbia, Canada
A portrait of a clown triggerfish (Balistoides conspicillum) on a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Misool Marine Protected Area. Ceram Sea. Tropical West Pacific Ocean
Tiger cardinalfish (Cheilodipterus macrodon) male mouth brooding a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea
A male pajama cardinalfish (Sphaeramia nematoptera) incubating eggs in his mouth (not clearly visible). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
British World War II BSA M20 motorbike inside Hold 2 of wreck of HMS Thistlegorm, home to pair of Red Sea soldierfish (Myripristis murdjan) and nudibranch (Nembrotha megalocera)
Detail of the filter feeding apendange of Peacock worm (Sabella pavonina) Loch Carron, Ross and Cromarty, Scotland, UK, April
Spotted eagle ray (Aetobatus narinari), Kicker Rock, San Cristobal Island, Galapagos