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Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
American bison (Bison bison) Yellowstone National Park, Montana, USA
Grizzly bear (Ursus arctos) catching a fish, Brooks Falls in Katmai National Park, Alaska, USA, July
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
European hare (Lepus europaeus), Wirral, England, UK, May
Jaguar (Panthera onca) male walking, Pantanal, Brazil
Arctic Fox (Alopex / Vulpes lagopus) standing on ridge, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Arctic Fox (Alopex / Vulpes lagopus) yawning, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Lions (Panthera leo) - two brothers patrolling territorial boundary, affectionate behaviour, border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Pine marten (Martes martes) reflected in water, Ardnamurchan Peninsula, west coast of Scotland, UK. Highly commended in the Mammals category of the British Wildlife Photography Awards (BWPA)
Weasel (Mustela nivalis) looking out of hole on woodland floor with snowdrops, Sheffield, England, UK
Young urban Red fox (Vulpes vulpes). Bristol, UK. August
Siberian tiger (Panthera tigris altaica) in snow, captive
Siberian musk deer (Moschus moschiferus) male in snow, Irkutsk, Russia. January
Sperm whale (Physeter macrocephalus) tail fluke above water as it dives below a rainbow, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species. Digital composite
African elephant (Loxodonta africana) herd walking in the plains during the dry season, Masai-Mara Game Reserve, Kenya. October
Male Saiga antelope (Saiga tatarica) in winter, The Black Lands National Park, Kalmykia, Russia
Red necked pademelon (Thylogale thetis) with joey peering from pouch, in scrubland at edges of rainforest, small macropod found in eastern Australia, Lamington National Park, Queensland, Australia
Cougar (Puma concolor) portrait, captive, occurs in Americas. Digitally manipulated
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair close-up Manuel Antonio National Park, Quepos, Costa Rica
RF - Lion (Panthera leo) male on savanna with dramatic storm clouds, thundery sky and light rays in the background. Ngorongoro Crater, Tanzania. Composite image
Eastern lowland gorilla (Gorilla beringei graueri) silverback named Chimanuka climbing a tree, Kahuzi-Biega National Park, South Kivu Province, Democratic Republic of Congo (DRC)
Bengal Tiger (Panthera tigris) six month old cub jumping on its mother, Bandhavgarh National Park, India
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Elephant (Loxodonta africana) herd walking in a line, Etosha National Park, Namibia
Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Adult Mountain gorilla (Gorilla beringei beringei) holding baby, Hirwa group, Volcanoes National Park, Rwanda
Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK. March
European Beaver (Castor fiber) eating blackberries by standing on its hind legs and using its tail as additional support. Part of captive breeding population prior to release into the Welsh landscape
Humpback whale (Megaptera novaeangliae) two breaching at the same time together, Puerto Lopez, Santa Elena Peninsula, Manabi Province, Ecuador, July, Puerto Lopez, Santa Elena Peninsula
Alpine ibex (Capra ibex) female, Jura, Switzerland, September
Meerkats (Suricata suricatta) with young, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
African wild dog (Lycaon pictus) interacting with pup age two months, Okavango Delta, Botswana
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa
Snow leopard (Panthera uncia) in snow, captive, USA
Bobcat (Lynx rufus) standing on branch in snow. Madison River Valley, Yellowstone National Park, Wyoming, USA. January
Brown bear (Ursus arctos) paw seem from under water, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August
RF- White horse of the Camargue, running from sea. Camargue, Southern France
Guanaco (Lama guanicoe), Punta Norte, Peninsula Valdes, Argentina
Cheetah (Acinonyx jubatus) staring back over its shoulder, black and white. Save Valley Conservancy, Zimbabwe, August
Red panda (Ailurus fulgens) captive, occurs in China
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
RF - Red Squirrel (Sciurus vulgaris) on log in snow. Scotland, UK. December
Timber Wolves (Canis lupus) resting on a rock by the shoreline. Near Mussel Inlet, Great Bear Rainforest, British Columbia, Canada, October
Pine marten (Martes martes) climbing Scots pine tree, Scotland, UK. May
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Lioness (Panthera leo) carrying her cub, Masai-Mara Game Reserve, Kenya
