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Grey wolf (Canis lupus) adult greeted by cub, captive, USA
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Pine marten (Martes martes) portrait, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Portrait category of the BWPA (British Wildlife Photography Awards) Competition 2016
Badger (Meles meles) adult, backlit at sunset, scratching its ears, Derbyshire, UK, May. Winner of the Animal Behaviour category of the British Wildlife Photography Awards (BWPA) Competition 2017
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
Blue whale (Balaenoptera musculus) underwater, Coronado Islands, Baja California, Mexico, Eastern Pacific Ocean
Impala with Oxpeckers. Kruger National Park, South Africa
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Giant panda (Ailuropoda melanoleuca) cub yawning. Yuan Meng, first giant panda ever born in France, age 10 months, Captive at Beauval Zoo, Saint Aignan sur Cher, France
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
Lion (Panthera leo) cubs playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Three Sperm whales (Physeter macrocephalus) socialising under the surface, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Gaur (Bos gaurus) Manas National Park UNESCO World Heritage Site, Assam, India
Red fox (Vulpes vulpes) in snowy landscape with trees and two Crows (Corfus corone) winter, Jura, Switzerland
Kiang (Equus kiang) herd grazing on steppe in evening light, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019
Sumatran tiger (Panthera tigris sondaica). Captive, with digitally added leaf pattern
Coyote (Canis latrans) walking through deep winter snow. Yellowstone National Park, Wyoming, USA. January
Red squirrel (Sciurus vulgaris) sitting on fallen tree in pine woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, August
American bison (Bison bison) male walking through deep snow. Hayden Valley. Yellowstone National Park, Wyoming, USA. January
Wild Eurasian lynx (Lynx lynx) at night with city lights and sky glow behind, Switzerland. December. Taken with remote camera trap
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Zanzibar red colobus monkey (Procolobus kirkii) crossing the main road through the Reserve, Jozani forest, Jozani Chwaka Bay NP, Zanzibar, Tanzania, August
Bearded pig (Sus barbatus) digging in sand, foraging for crabs on beach, Bako National Park, Sarawak, Borneo
Grey whale (Eschrichtius robustus) tail, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Red squirrel (Sciurus vulgaris) gnawing red deer antler for minerals, Cairngorms National Park, Highlands, Scotland, UK, June
Red squirrels (Sciurus vulgaris) interacting, Cairngorms National Park, Highlands, Scotland, UK, January
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
Badger (Meles meles) cub amongst long grass, Dorset, England, UK, July
Red fox (Vulpes vulpes) resting in the snow, Churchill, Cananda, November
Red squirrel (Sciurus vulgaris) having a drink, Black Isle, Scotland, UK. October
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Desmarests hutia (Capromys pilorides doceleguas), Jardines de la Reina / Gardens of the Queen, Caribbean Sea, Ciego de Avila, Cuba, January
Two Sulawesi black macaques / Celebes crested macaques (Macaca nigra) juveniles, in woodland, portrait, Tangkoko National Park, northern Sulawesi, Indonesia. Critically endangered
Australian sea (Neophoca cinerea) juvenile playing in the sand, Carnac Island, Western Australia
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Barbary macaque (Macaca sylvanus), two adults with baby, overlooking town and ships in sea at sunset. Gibraltar Nature Reserve, Gibraltar. August 2018
Bottlenose dolphins (Tursiops truncatus) at the surface at sunrise. Kwazulu-Natal, South Africa
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
Brown bear (Ursus arctos arctos) female and two cubs, aged 1 year, emerging from den in winter, Bavarian Forest, Germany, captive, January
1468136 - Mandrill male (Mandrillus sphinx) abstract of brightly coloured ischial1468136 - - Mandrill male (Mandrillus sphinx) abstract of brightly coloured ischial callosity on buttocks, Lekedi National park, Gabon
Aerial view of Reindeer (Rangifer tarandus) fording water whilst being herded by Nenet herders. Nenets Autonomous Okrug, Arctic, Russia, July 2017
Grey Seal (Halichoerus grypus) at rest on submerged rock, head and one flipper above water. Shetland Islands, Scotland, UK. July
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair playing with each other, one with hand over eyes of other, Manuel Antonio National Park, Quepos, Costa Rica
Black rhino (Diceros bicornis) and White Rhino (Ceratotherium simum) bulls facing off. Solio Game Reserve, Laikipia, Kenya. September
