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Indian wild ass (Equus hemionus khur) running, Wild Ass Sanctuary, Little Rann of Kutch, Gujarat, India
Golden snub-nosed monkey (Rhinopithecus roxellana), mother and baby, Qinling Mountains, Shaanxi province, China
Arctic fox cub (Alopex lagopus) portrait in grass with summer flowers, Hornvik, Hornstrandir, Westfjords, Iceland. July
RF - Arctic fox cub (Alopex lagopus) at den. Hornvik, Hornstrandir, Westfjords, Iceland. July. (This image may be licensed either as rights managed or royalty free.)
Altai argali sheep (Ovis ammon), three grazing on slope. Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China. October
Tibetan gazelle (Procapra picticaudata) in snow. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October
RF-Asian elephant (Elephas maximus) dust bathing. Jim Corbett National Park, Uttarakhand, India. (This image may be licensed either as rights managed or royalty free)
Pallass cat (Otocolobus manul) licking lips whilst running over ice. East Mongolia. February
Snow leopard (Panthera uncia) with ears back. Captive
Eurasian lynx (Lynx lynx) yawning. Captive
Lion (Panthera leo) males mock fighting / play fighting. Ngorongoro Crater, Ngorongoro Conservation Area (NCA), Tanzania
Red fox (Vulpes vulpes) running through snow. Hayden Valley, Yellowstone, USA. January
Fosa / Fossa (Cryptoprocta ferox) male resting in dry decidous forest. Kirindy, western Madagascar. Endangered
Southern right whale (Eubalaena australis) diving, with tail fluke splash. Monumento Natural Ballena Franca Austral, UNESCO World Heritage Site, Valdes Peninsula, Patagonia, Argentina
Sunda clouded leopard (Neofelis diardi) at night, Deramakot Forest Reserve, Sabah, Malaysian Borneo
Moose bull (Alces alces) walking in forest clearing, Denali National Park, Alaska, USA, September
Javan leopard (Panthera pardus melas) roaring, native to the Indonesian island of Java. Captive. Digital composite
Indri (Indri indri) with young, Palmarium Resrve Ankanin ny Nofy, Littoral rainforest, Madagascar
Golden jackal (Canis aureus) pup sitting on a grassy verge by a path, Danube Delta, Romania, July
African lion (Panthera leo) and lioness after mating, Masai Mara National Reserve, Kenya
Lion (Panthera leo) male walking. Masai Mara National Reserve, Kenya
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Grey-headed flying-fox (Pteropus poliocephalus) spotting and flaring wings to come in to land on a branch. Yarra Bend Park, Kew, Victoria, Australia. ÔÇ¿November 2019
Fosa of Fossa (Cryptoprocta ferox) male walking through dry deciduous forest, Kirindy, western Madagascar
Aye-aye (Daubentonia madagascariensis) adult active and foraging in forest canopy at night. Deciduous forests, Daraina, northern Madagascar. Endangered endemic species
Red fox (Vulpes vulpes) adult hunting for rodents by snow diving in deep snow. Hayden Valley, Yellowstone, USA. January
Boky-boky (Mungotictis decemlineata) or Narrow-striped mongoose adult foraging in leaf litter. Dry deciduous forest, Kirindy, western Madagascar
Fosa or Fossa (Cryptoprocta ferox)(sometimes erroneously fossa) male in dry deciduous forest. Kirindy, western Madagascar
Puma (Puma concolor puma), female sitting in grassland. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. November
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Proboscis monkey (Nasalis larvatus) Riverine forest, Kinabatangan River, Sukau, Sabah, Borneo
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Giant River Otter (Pteronura brasiliensis) feeding on Striped Catfish or Cachara (Pseudoplatystoma fasciatum) amongst water hyacinth. Tributary off the Cuiaba River, Mato Grosso, Pantanal, Brazil
Siberian flying squirrel (Pteromys volans orii) feeding on Sakhalin fir tree. Hokkaido, Japan. March
North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching at sunset, Kupreanof Island, Frederick Sound, Alaska, USA, July
Killer whale / Orca (Orcinus orca) mature male, swimming underwater. Hamn, Senja, Norway, Atlantic Ocean
Humpback whale calf (Megaptera novaeangliae) male resting beneath his resting mother. Vava u, Tonga, South Pacific
Atlantic walrus (Odobenus rosmarus rosmarus) hauled out on ice floe, Svalbard, Norway June
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Giant panda (Ailuropoda melanoleuca) cub in tree, Chengdu Panda Breeding Centre, Sichuan, China. Captive
Short-finned pilot whale (Globicephala macrorhynchus) calf with fetal lines characteristic of newborn specimens, with pod. Tenerife, Canary Islands
Arctic fox (Alopex lagopus) camouflaged in winter pelage. Svalbard, Norway, April
Polar bear (Ursus maritimus) standing on ice. Svalbard, Norway, July
Roe Deer stag (Capreolus capreolus) standing in a woodland glade. Potterick Carr Nature Reserve, Doncaster, Yorkshire, England, UK, July
Lion (Panthera leo) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa