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Wild boar (Sus scrofa) camera trap image in the Jura mountains, Switzerland, September
Collared peccary (Pecari tajacu) Laredo Borderlands, Texas, USA. April
Warthogs (Phacochoerus aethiopicus) mother with babies drinking, Mkhuze game reserve, KwaZuiu-Natal, South Africa, June
Warthog (Phacochoerus africanus) mother with baby on back sleeping, Masai Mara, Kenya
Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night
Wild boar (Sus scrofa) piglets (known as humbugs ) in woodland clearing. Forest of Dean, Gloucestershire, UK. May
Wild boar (Sus scrofa) in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
Golden jackal (Canis aureus) portrait. Danube Delta, Romania, May
Gurskys spectral tarsier (Tarsius spectrumgurskyae) North Sulawesi
Horsfields tarsier / Western tarsier ( Tarsius bancanus ssp. saltator) Belitung Island, Sumatra, Indonesia. Endemic
South Georgia pintail (Anas georgica georgica) swimming in front of Southern elephant seal (Mirounga leonina), male. Gold Harbour, South Georgia. October
African wild dog (Lycaon pictus) portrait, Mana Pools National Park, Zimbabwe
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
Iberian lynx (Lynx pardinus) with Magpie (Pica pica) behind, Sierra de Andujar Natural Park, Jaen, Spain, September
Grey whale (Eschrichtius robustus) eye, Magdalena Bay, Baja California, Mexico, February. Winner in the Ocean Views category of the 2018 Natures Best Winland Smith Rice Awards
Red fox (Vulpes vulpes) adult with train in background, Kent, UK. March. Highly commended in the Nature Photographer of the Year Awards 2018. Taken with remote camera
White rhinoceros (Ceratotherium simum) five calves, orphaned from poaching feeding from a trough at dusk at the Rhino Revolution orphanage near Hoedspruit, South Africa. May 2017
Scarred face of a white rhinoceros (Ceratotherum simum) that survived an attack by poachers who illegally tranquillised her, two other rhinoceroses died from their injures
Aerial view of a herd of wild White rhinoceros (Ceratotherium simum) running free on Chiefs Island, Okavango Delta, Botswana
Young White rhinoceros (Ceratotherium simum) in a reinforced steel boma, blindfolded and partially drugged after a long journey from South Africa, before being released into the wild in Botswana
Mountain hare (Lepus timidus) in extreme winds with spin drifts of snow, Cairngorms National Park, Scotland, UK, February 2018
Pallass cat (Otocolobus manul) Tibetan Plateau, Qinghai, China
Leopard (Panthera pardus) cub looking up at birds (out of frame) with mother in background, Khwai River, Botswana
Black spider monkey (Ateles chamek) calling, captive, Peru
White lipped peccary (Tayassu pecari) on forest floor, Lago Preto Conservation Concession, Yavari River, Peru
Black spider monkey (Ateles chamek) portrait, captive, Peru
Golden-mantled tamarin (Saguinus tripartitus) captive
Drill monkey (Mandrillus leucophaeus) adult male, portrait, captive
Tibetan macaque (Macaca thibetana) Tangjiahe Nature Reserve, Sichuan Province, China
Indo-Pacific humpback dolphin (Sousa chinensis) with fishing net damage around throat, Pearl River Estuary, Lantau Island, Hong Kong, China
RF - White lion (Panthera leo) male, portrait of head. Captive, Netherlands
Springbok (Antidorcas marsupialis) herd fleeing predator, Kgalagadi Transfrontier Park, South Africa
RF - Meerkat (Suricata suricatta) resting on cool sand, Kgalagadi Transfrontier Park, South Africa
Straw-coloured fruit bat (Eidolon helvum), female flying carrying pup on front. Lamin, Gambia
Jaguar (Panthera onca) male, hunting Capybara (Hydrochoerus hydrochaeris). The capybara while jumping away has cut the nose of the Jaguar with its toemail
Jaguar (Panthera onca) male, chasing a Caiman. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
African lion (Panthera leo) growling at potential danger in the water (Panthera leo) Okavango Delta, Botswana
Male Lion (Panthera leo) portrait, close-up of face, captive, occurs in Africa
Lioness (Panthera leo) resting up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
Lion (Panthera leo) cub up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
Lion (Panthera leo) male standing in cold and rain with strong wind blowing, smelling the air, Masai Mara National Reserve, Kenya
Two male African lions (Panthera leo) drinking at a waterhole, Sabi Sand Private Game Reserve, South Africa
Male Lion (Panthera leo) Romeo in habitat, from the Marsh Pride, Masai Mara, Kenya
African lion (Panthera leo) three subadults resting on the road, one snarling, Central Kalahari Game Reserve, Botswana
Female Bornean orangutan (Pongo pygmaeus) sitting in a tree, Tanjung Puting National Park, Borneo-Kalimantan, Indonesia
Black rhino (Diceros bicornis) female browsing on herbs. Ngorongoro Crater, Conservation Area, Tanzania. March
Black rhinoceros (Diceros bicornis) covered in mud, Mud on skin Etosha National Park, Namibia
Jaguar (Panthera onca) mother grooming with four month old cub, native to Southern and Central America, captive
