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Portrait of a Blackbar soldierfish (Myripristis jacobus) infected with Cymothoid isopod parasites. The large parasite between the eyes is a female, the smaller individuals are males
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
Azure / common coenagrion damselflies (Coenagrion puella) pair mating, Meeth Quarry, Devon, UK. July
Portrait of two siika deer (Cervus nippon), possibly red deer (Cervus elaphus) hybrids, on open moorland in snow, Cairngorms National Park, Scotland, UK. January 2020
Common shelduck pair (Tadorna tadorna) standing and drinking in the margins of a shallow lake, Gloucestershire, UK, November
Mute swan (Cygnus olor) pair preening in unison, Gloucestershire, UK, December
Galapagos hawk (Buteo galapagoensis) landing on mating pair of Galapagos giant tortoise (Chelonoidis vandenburgh) Alcedo Volcano, Isabela Island, Galapagos
Waved albatrosses (Phoebastria irrorata) courting, Punta Suarez, Espanola Island, Galapagos
Swallow-tailed gull (Creagrus furcatus) pair mutual preening in courtship. Genovesa Island, Galapagos Islands, Ecuador, June
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Northern Cardinals (Cardinalis cardinalis) male and female in the snow, photographed from low angle, Freeville, New York, USA. April 2020
Snow leopard (Uncia uncia) pair resting in sunshine amongst rocks. Altai Mountains, West Mongolia. February
RF-Snow leopard (Uncia uncia) pair sitting on ledge of rockface. Altai Mountains, West Mongolia. February. (This image may be licensed either as rights managed or royalty free)
Leopard (Panthera pardus) pair mating behaviour, Okavango Delta, Botswana
Mute swan (Cygnus olor) pair at sunrise. London, UK. December
Pair of courting Egg-eating sea snakes / Turtleheaded sea snakes (Emydocephalus annulatus) New Caledonia, Pacific Ocean
Eurasian rollers (Coracias garrulus) courting on branch, Hungary. June
Small emerald spreadwing (Lestes virens) pair mating. Klein Schietveld, Brasschaat, Belgium. August
Blue-tailed damselfly (Ischnura elegans) pair mating on stem. Brasschaat, Belgium. August
King cormorants (Phalacrocorax atriceps) in courtship behaviour at their nest site. Saunders Island, Falkland Islands
Polar bear (Ursus maritimus) male female pair looking upwards, snow covered hills in background. Svalbard, Norway, April 2018
Polar bear (Ursus maritimus) male approaching female in rocky area covered in snow. Svalbard, Norway, April 2019
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Hyacinth macaw (Anodorhynchus hyacinthinus) pair perched on branch, silhouetted under crescent moon. Mato Grosso Forest, Pantanal, Mato Grosso, Brazil
Amur / Siberian tiger (Panthera tigris altaica) pair in courtship, in snow. Captive in tiger park, Heilongjiang Province, China. January
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef
Moose (Alces alces) female and male covered in light dusting of snow in forest, Grand Teton NP, Wyoming, USA, October
Muskox {Ovibos moschatus} pair on the tundra in autumn, Dovrefjell National Park, Norway, September 2008
Nurse sharks (Ginglymostoma cirratum) three in a courtship dance at sunrise in a mangrove area near Eleuthera, Bahamas
Fairy Tern (Gygis alba) pair, Cousin Island, Republic of Seychelles
Laysan albatross (Phoebastria immutabilis), courtship ritual, Eastern island, Midway Atoll National Wildlife Refuge, Hawaii
Red Footed Booby (Sula sula) couple, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Common toad (Bufo bufo) pair in amplexus, Deeside, Scotland April
Iolas blue butterfly (Iolana iolas) pair mating on flowers, Aosta Valley, Gran Paradiso National Park, Italy
Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Veragua cross-banded treefrogs (Smilisca sordida) mating pair at night, Osa Peninsula, Costa Rica
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
Velvet Swimming Crab (Necora / Liocarcinus puber) male guarding female. Gouliot Caves, Sark, British Channel Islands, August
Razorbill (Alca torda), pair on cliff, Farne Islands, Northumberland, England, UK, June
Northern fulmar (Fulmarus glacialis) nesting pair, Saltee Islands, County Wexford, Republic of Ireland, June
Northern gannets {Morus bassanus} pair in courtship display, Great Saltee, Co. Wexford, Ireland, June
African lion (Panthera leo) male female pair reflected in water, Okavango Delta, Botswana
Orange tip butterfly (Anthocharis cardamines) female and male, visiting Honesty flower (Lunaria annua) Wiltshire, England, UK, May
Diademed sifaka (Propithecus diadema diadema) pair, Maromizaha Reserve, Andasibe Mantadia area, eastern Madagascar. August
Horned puffins (Fratercula corniculata) pair interacting by touching bills while perched on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Mating pair of Azure damselflies (Coenagrion puella) on Plantain (Plantago sp) Lagadin region, Lake Ohrid, Galicica National Park, Macedonia, June 2009
Alpine grasshopper (Miramella alpina) pair mating on a thistle, Triglav National Park, Slovenia, August 2009
Robber flies (Asilidae) mating, one with insect prey, The Peloponnese, Greece, May 2009
