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Red-collared lorikeet (Trichoglossus rubritorquis) pair examining hole in Eucalyptus as a potential nesting hollow. Katherine, Northern Territory, Australia
Kestrel (Falco tinnunculus) fighting Hungary, April
Northern raven (Corvus corax) pair perching on branch. Danube Delta, Romania, May
White-tailed eagle (Haliaeetus albicilla) male and female perched on tree in snow, near Kuusamo, Finland. March. Runner up in the Birds category of the GDT European Wildlife Photographer of the Year
African lion (Panthera leo) pair, Masai Mara National Reserve, Kenya, January
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Wood pigeon (Columba palumbus) pair preening one another, The Netherlands
RF - Southern elephant seal (Mirounga leonina) male and female, Valdes, Patagonia Argentina, Word Heritage
Tawny nurse shark (Nebrius ferrugineus) group, individuals circling each other at dusk. Vavuu Atoll, Maldives. Indian Ocean
BlackbuckA(Antilope cervicapra) herd with males and females, Velavadar national parkBlackbuck(Antilope cervicapra) herd with males and females, Velavadar national park, Gujarat, India
Silhouette of Peregrine falcon (Falco peregrinus) in flight, male passing pigeon prey to female for her to feed the chicks, Barcelona, Spain, April
Greater flamingo (Phoenicopterus roseus) pair mating, in lagoon, Pont Du Gau, Camargue, France, April 2009
Pair of mating soldier beetles (Rhagonycha fulva) on Common ragwort (Senecio jacobaea), Arne RSPB reserve, Dorset, England, UK, August
European Adder (Vipera berus) portrait of a female and a male. Aberdeenshire, Scotland, May
Red deer (Cervus elaphus) stag and hind at dusk, Cheshire, UK September
Palmate newt (Triturus helveticus) male and female underwater, captive
Kingfisher (Alcedo atthis) male passing fish to female spring courtship behaviour, Hertfordshire, England, UK, March. Sequence 4 of 6
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Small emperor moth (Saturnia pavonia) male below female both displaying eyespots on Gorse, Captive, UK, April
Male Carolas Parotia (Parotia carolae) bird of paradise displaying on his display court to five females perched above, Crater Mountain Wildlife Management Area, Eastern Highlands Province
Waved Albatross (Phoebastria irrorata) courtship, Punta Cevallos, Espaola Island, Galapagos Islands, Ecuador, South America. Endemic, Critically endangered
Scarlet macaw (Ara macao) pair in tree, Osa Peninsula, Costa Rica
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Mute swan (Cygnus olor) pair courting. Walthamstow reservoir, London, UK
Rio pescado harlequin toad (Atelopus balios) pair on branch, female on the left, Azuay, Ecuador, Critically endangered species, possibly extinct
Jackdaw (Corvus monedula) pair chasing a Grey squirrel (Sciurus carolinensis) as it emerges from a nest box the birds want to nest in which is already occupied by the squirrel and its mate
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Kingfisher ( Alcedo atthis) male and female perched on a root when building the nest, Lorraine, France, March
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
Black-veined white (Aporia crataegi) butterfly pair, in flight and nectaring on Thistle. Jyvaskyla, Central Finland. June
Espanola saddelback tortoise (Chelonoidis hoodensis) pair mating, Espanola Island, Galapagos
Snow leopard (Uncia uncia) pair on snow covered rocky slope. Altai Mountains, West Mongolia. February
Pair of courting Egg-eating sea snakes / Turtleheaded sea snakes (Emydocephalus annulatus) New Caledonia, Pacific Ocean
Black Woodpecker (Dryocopus martius) male and female, at the nesthole, Germany. December
Colorado river toad (Incilius alvarius) pair mating. Sonoran desert, Arizona, USA. August
Manchurian crane (Grus japonensis) pair in courtship dance during snowstorm. Hokkaido, Japan. March
Greater flamingo (Phoenicopterus roseus) pair at rest in water, Cape Town, South Africa, July
Grey seal (Halichoerus grypus), male and female, mating behaviour, Heligoland, Germany
Golden pheasant (Chrysolophus pictus) male and female, Yangxian nature reserve, Shaanxi, China
Western clown anemonefish (Amphiprion ocellaris) poses with a large, dominant female behind. The clownfish in the foreground is probably a juvenile. Bitung, North Sulawesi, Indonesia
Scarab beetle (Scarabaeoidea) pair mating inside Tulip (Tulipa micheliana) flower. Beetles covered in pollen. Uzbekistan, April
Bumblebee scarab (Amphicoma vulpes) pair mating in Tulip (Tulipa armena) flower. Pollen picked up on beetles and transferred to other flowers. Near Karabel Pass, Turkey. May
Amur / Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snowAmur/ Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snow. Captive in tiger park, Heilongjiang Province, China. February
Rhinoceros hornbills (Buceros rhinoceros) pair flying over the canopy with mist hanging over lowland Dipterocarp rainforest in heart of Danum Valley, Sabah, Borneo (digitally modified image)
Black howler (Alouatta caraya) male and female, captive, occurs in Brazil and Paraguay. With digitally added leaf pattern
