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Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia
Western lowland gorilla (Gorilla gorilla gorilla) male aged 12 years standing up at edge of Mpassa river. Captive bred in UK and released into wild along with his father in 2013
Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. June
Andean condor (Vultur gryphus), Male, Antisanilla Reserve, Ecuador
Okarito Brown Kiwi (Apteryx rowi) male known as Scooter patrolling his territory, from population of 200. Okarito Forest, Westland, South Island, New Zealand
Tiger (Panthera tigris tigris) dominant male tiger approaching Beja pani waterhole, Kanha National Park, Central India. Camera trap image
Indian Flying Fox (Pteropus giganticus) male roosting in tree, Keoladeo NP, Bharatpur, India
Orca (Orcinus orca) surfacing and blowing, Shetland, Scotland, UK, August
Roe deer (Capreolus capreolus) buck with vegetation on antlers in rutting season, Scotland, UK. August
Tufted duck (Aythya fuligula) male drake, Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Mandarin duck drake (Aix galericulata) on the water, showing orange sail feathers which are used when displaying, Southwest London, UK, March
Banded demoiselle (Calopteryx splendens) male, resting on grass head, Lower Tamar Lakes, Cornwall, UK. June
Cinnamon teal (Anas cyanoptera), male taking flight from water, Orange County, California, USA, April
California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Eastern bluebird (Sialia sialis) male fluttering wings while perched on cherry (Prunus sp.) branch in spring, New York, USA, April
Bengal male cat stalking
Running foliage spider / Spiny-legged sac spider (Liocranum rupicola), a nationally scarce species in the UK, hunting at night in an old stone wall, Wiltshire, UK, September
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Cuckoo (Cuculus canorus) male, Germany, April
Blackcap (Sylviya atricapilla) male perched in blossom, Hungary, April
Great bowerbird (Chlamydera nuchalis), male showing off treasures whilst female inspects bower. Lake Argyle, Kununurra, Western Australia
Tapanuli orangutan (Pongo tapanuliensis) Togus, adult flanged male, Batang Toru Forest. Sumatran Orangutan Conservation Project, North Sumatran Province, Indonesia
Blackbird (Turdus merula) in flight to feed on berries, Helsinki, Finland. November
Bearded reedling (Panurus biarmicus) on frosty reed seed head, Helsinki, Finland, January
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
Male Golden Pheasant (Chrysolophus pictus) standing on grass in display plumage. Kew Gardens, England, UK. March
Pine grosbeak (Pinicola enucleator), male feeding juvenile, Finland, July
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Long-tailed duck (Clangula hyemalis), males in flight, Finland, April
Black faced sheep ram with twisted horns, Mull, Scotland, UK. January
Norfolk Terrier dog, Otto, age 1 year
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Chesapeake Bay retriever male standing in saltmarsh, Long Island Sound, Madison, Connecticut, USA
Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
South Georgia pintail (Anas georgica georgica) swimming in front of Southern elephant seal (Mirounga leonina), male. Gold Harbour, South Georgia. October
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
Stag beetle (Lucanus cervus) males fighting on oak tree branch, Elbe, Germany, June
Drill monkey (Mandrillus leucophaeus) adult male, portrait, captive
American moon moth (Actias luna) male, North America
Northern crested newt (Triturus cristatus) male underwater in a pond, during the mating season. Isere, Cremieu, France
RF - White lion (Panthera leo) male, portrait of head. Captive, Netherlands
African lion (Panthera leo) growling at potential danger in the water (Panthera leo) Okavango Delta, Botswana
Male Lion (Panthera leo) portrait, close-up of face, captive, occurs in Africa
Lion (Panthera leo) male standing in cold and rain with strong wind blowing, smelling the air, Masai Mara National Reserve, Kenya