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Short eared owl (Asio flammeus) tilting head upside down and backwards, Breton Marsh, West France, June
Two young Polar bears (Ursus maritimus) on newly formed pack ice, near Kaktovik, Barter Island, North Slope, Alaska, USA, October. Vulnerable species
Tabby kitten, Picasso, age 3 months, sleeping on a fluffy rug
Tabby kitten, Picasso, 8 weeks, with raised paw
Eurasian otter (Lutra lutra) portrait, Scotland, UK, April
Whooper swan (Cygnus cygnus) pair on loch in heart shape, Cairngorms National Park, Scotland, UK, November
Black throated diver (Gavia arctica) swimming, Assynt, Sutherland, Highland, Scotland, UK, June
View to The Quiraing from the Trotternish Ridge at dawn, Isle of Skye, Inner Hebrides, Scotland, UK, April 2016
European river otter (Lutra lutra) female bringing octopus ashore for cub, Shetland, Scotland, UK, October
A family of otters rest on the intertidal seaweed. European river otter (Lutra lutra) Shetland, Scotland, UK, July
Black bears (Ursus americanus) three cubs, one climbing, Minnesota, USA, June
Ferruginous Pygmy-Owl (Glaucidium brasilianum). Yucatan. Mexico. February
Fledgling Northern Hawk Owl (Surnia ulula). Finland. June
Fer-de-lance (Bothrops asper) camouflaged on the rainforest floor. Corcovado National Park, Osa Peninsula, Costa Rica
Fleischmanns glassfrog (Hyalinobatrachium fleischmanni) male attending eggs on the underside of a leaf overhanging a rainforest stream
Jumping spider (Salticidae) hunting among vegetation, San Jose, Costa Rica
Granular poison frog (Oophaga granulifera) Osa Peninsula, Costa Rica. Vulnerable IUCN Red List species
Apollo butterfly (Parnassius apollo) Nordtirol, Austrian Alps. June
Nine-spotted moth (Amata phegea) feeding from a Mountain arnica flower (Arnica montana) Triglav National Park, Julian Alps, Slovenia. July
Painted lady butterfly (Vanessa cardui) on flower, Peak District National Park, Derbyshire, UK. September
Red Squirrel (Sciurus vulgaris) leaping onto log, Scotland, UK, September. Highly commended in the Habitat category of the BWPA (British Wildlife Photography Awards) 2016
Grass snake (Natrix natrix) flicking tongue, Hampstead Heath, London, England, UK, August
Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea
California sea lions (Zalophus californianus) group playing in the sun in the early morning. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Southern stingrays (Hypanus americanus) swimming over a sand bar in the early morning. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Portrait of a male Squarespot anthias (Pseudanthias pleurotaenia). Sangeang Island; Sumbawa, Indonesia. Flores Sea
Brandts cormorant (Phalacrocorax penicillatus) swimming beneath an oil rig. Eureka Rig, Los Angeles, California, United States of America. North East Pacific Ocean
A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
Great white sharks (Carcharodon carcharias) males beneath the surface. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Green-winged orchids (Anacamptis morio) at sunrise, Ashton Court Park, Bristol, UK, May
Yellow tube sponge (Aplysina fistularis) on a coral reef, with the camera rotated during the long exposure. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Black skimmer (Rynchops flavirostris) fishing by skimming along water surface, Pantanal, Brazil
Loch Beinn a Meadhoin in the mist, Inverness, Scotland, December 2013
Lava Lake at night in the crater of Nyiragongo Volcano, Virunga National Park, North Kivu Province, Democratic Republic of Congo. September 2015
Indri (Indri indri) feeding on fresh leaves / shoots in the rainforest canopy. Andasibe-Mantadia National Park, eastern Madagascar
Cup fungus (Cookeina sp) growing on decaying wood on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Tranquil landscape with African elephant (Loxodonta africana) and rays of sunlight at sunrise, Masai Mara, Kenya
Masai giraffe (Giraffa camelopardalis tippelskirchi) two silhouetted at sunrise, Msai Mara Game Reserve, Kenya
Runner on sunlit path in early morning, cose to the Vale of Health Hampstead Heath, London, England, UK. November 2015
Water cascading over naturally formed tufa dam. Tufa is a kind of limestone, formed from the precipitation of carbonate minerals which builds up around mosses
Mute swan (Cygnus olor) cygnets on water, Lac de Saint Point, Franche Comte, France, May
Red fox (Vulpes vulpes) cub biting tail of adult male, Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East. June
Graylag gosling (Anser anser) with parent, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. April
White rhinoceros (Ceratotherium simum) charging, Mala Mala Game Reserve, South Africa
Kirkjufellsfoss waterfall at sunrise, with Kirkjufell mountain in the background, Snaefellsnes Peninsula, Western Iceland June
Amethyst Deceiver (Laccaria amethystina) two young toadstools growing up from mossy log, Buckinghamshire, England, UK, September
Kaieteur Falls is the worlds widest single drop waterfall, located on the Potaro river in the Kaieteur National Park, in Essequibo, Guyana, South America