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Glorious scarab (Chrysina gloriosa) flying, Brewster County, Texas, USA. Controlled conditions. August
Leaf cutter ants (Atta sp) carrying pieces of fern, reflected in water, Costa Rica
Hyacinth macaws (Anodorhynchus hyacinthinus) mutual preening, Estrada Parque, Pantanal, Brazil
Green sea turtle female (Chelonia mydas) returning to the sea, Tortuguero National Park, Costa Rica
Green sea turtle (Chelonia mydas) female resting on a coral reef. Rock Islands, Palau, Mirconesia. Tropical west Pacific Ocean
Hercules moth (Coscinocera hercules) recently emerged in montane rinforest. Ambua Lodge, Tari, Hela Province, Papua New Guinea. June
Queen Marys Tree, a veteran Sweet chestnut (Castanea sativa) over 600 years old, Cumbernauld, Scotland, UK, July
Full moon rising over the Verpeilspitze (3430m). This peak is part of the Glockturmkamm, the westernmost ridge of the Otztal Alps. Nordtirol, Austrian Alps. July. Digitally stitched panorama
Canadian Rockies reflected in marshland, Banff National Park, Alberta, Canada
California sea lions (Zalophus californianus) two resting with flippers round each other, Monterey Bay, California, USA, September
Reticulated giraffe (Giraffa camelopardalis reticulata) mother grooming baby, Samburu Game Reserve, Kenya, Africa, August
Mountain gorillas (Gorilla beringei beringei) baby sticking out tongue, Volcanoes National Park / Parc National des Volcans, Rwanda
African elephants (Loxodonta africana) swimming across Chobe River, trunks raised above surface for breathing, Chobe National Park, border between Botswana and Namibia
Humpback whale (Megaptera novaeangliae), fluke of female with male calf playing below, Vava'u, Tonga, Pacific Ocean
Ancient Beech trees (Fagus sylvatica), Lineover Wood, Gloucestershire UK. The second largest Beech tree in England in the foreground. November 2015
Gemsbok (Oryx gazella) aerial view of herd on sand, Namib Desert, Namibia
European bee eaters (Merops apiaster) one taking off another perched, Kiskunsagi National Park, Pusztaszer, Hungary. May
Monastery terraces on Skellig Michael, Skellig Islands World Heritage Site, County Kerry, Ireland, Europe. September 2015
Magnificent Bird of Paradise (Diphyllodes magnificus) male, Lower montane highlands near Mount Hagen, Enga Province, Papua New Guinea. Endemic
Waglers pit viper (Tropidolaemus wagleri) basking in mist-shrouded forest under storey. Danum Valley, Sabah, Borneo
Atlantic walruses (Odobenus rosmarus rosmarus) hanging out in shallow water in Svalbard, , Norway, June
Polar bear (Ursus maritimus) female with a single young cub, only a few months old, northern Svalbard, Norway, June
Blue whale (Balaenoptera musculus) swimming ins motion, Sri Lanka, Indian Ocean
Leaf cutter ants (Atta sp) carrying piece of leaf, Costa Rica
Alpine ibex (Capra ibex) on a mountain side in autumn with European larch tree (Larix decidua) trees in the back the Alps, Gran Paradiso National Park, Italy. November
Red deer (Cervus elaphus) male stag silhouetted at dusk, rut season, Jura, Scotland, UK, September
Water vole (Arvicola amphibius) Kent, UK September
Grey seal (Halichoerus grypus) playful young seal above a kelp forest. Lundy Island, Devon, UK Bristol Channel, August
Tricolour merle Collie puppy, Indie, age 10 weeks
Sperm whale (Physeter macrocephalus) with mouth open, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Blue lupin (Lupinus angustifolius) Sierra de Grazalema Natural Park, southern Spain, May
Red deer (Cervus elaphus) stag and two hinds, Los Alcornocales Natural Park, southern Spain, September
Lions (Panthera leo) five drinking together, Ndutu, Tanzania
Lion (Panthera leo) photographed at night using a side lit spot light, Sabi Sand Game Reserve, South Africa
Lioness (Panthera leo) sitting during misty sunrise, Okavango Delta, Botswana
Lionesses (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa. Sequence 5 of 5
Arctic fox (Alopex lagopus) juveniles play fighting, Wrangel Island, Far East Russia, August
Northern Cardinal (Cardinalis cardinalis), adult male perched on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Japanese / Red-crowned crane (Grus japonicus) one coming into land, Hokkaido Japan February
Highland cow in snow, Glenfeshie, Cairngorms National Park, Scotland, UK, February
Asiatic elephant (Elephas maximus), herd drinking water and crossing Mountain River at dawn, Jim Corbett National Park, India
Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic
Northern tamandua (Tamandua mexicana) Nicoya Peninsula, Costa Rica
Leaf mantis (Choeradodis sp. ) Tortuguero National Park, Costa RicaLeaf mantis (Choeradodis sp.) Tortuguero National Park, Costa Rica
Neotropical river otter (Lontra longicaudis) in water, Nicoya Peninsula, Costa Rica, March 2015
Great egret (Ardea alba) in snow, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. March
Short eared owl (Asio flammeus) tilting head upside down and backwards, Breton Marsh, West France, June
Two young Polar bears (Ursus maritimus) on newly formed pack ice, near Kaktovik, Barter Island, North Slope, Alaska, USA, October. Vulnerable species