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Great egret (Ardea alba) in flight, Florida, USA, March
Ice on black volcanic sand beach with waves, Jokulsarlon, Southeast Iceland, February
Trevose Lighthouse at dusk, long exposure at high tide, Trevose, Cornwall, UK. January 2015
Whooper swan (Cygnus cygnus) in flight in snowfall, Lake Kussharo, Japan, February
Snow Monkey (Macaca fuscata) with snow covered paw in front of mouth, Nagano, Japan, February
Coastal brown bear (Ursus arctos) resting, Lake Clarke National Park, Alaska, September
Dallol hot spring with salt concretions coloured by sulphur, potassium and iron, Dallol Volcano, Danakil Depression, Ethiopia, March 2015
Maple leaves backlit on lightbox, Broxwater, Cornwall, UK. November 2013
Green-veined White butterfly egg (Pieris napi) above, Orange-tip butterfly egg (Anthocharis)Green-veined White butterfly egg (Pieris napi) above, Orange-tip butterfly egg (Anthocharis cardamines) below, on Garlic mustard (Alliaria petiolata) Sussex, UK. June
Four Yellow-eyed penguins (Megadyptes antipodes) walking up a sand dune towards their nests, Otago Peninsula, Otago, South Island, New Zealand. September. Endangered Species
Selfoss waterfall in autumn with snow, north Iceland, September 2013
Takob-Akob Falls plunging 38 metres through the rainforest. Southern plateau edge, Maliau Basin, Sabah, Borneo, May 2011
Bearded pig (Sus barbatus) foraging for food at night, Maliau Basin, Sabah, Borneo
Two male Thomsons gazelles (Eudorcas thomsonii) fighting, Masai-Mara game reserve, Kenya
Ring tailed lemur (Lemur catta) mother and very young (1-2 week) baby. Berenty Private Reserve, Madagascar
Emperor penguin (Aptenodytes forsteri) chicks, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November