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Stars and Milky Way over Durdle Door and the Jurassic Coast, with the lights of Weymouth and Portland beyond, Dorset, England, UK, December 2013
Cape buffalo (Syncerus caffer) crossing the Chobe River, followed by swarm of flies, Chobe Game Reserve, Botswana, Africa
Great white egret (Ardea alba) splashing, Hungary, January
Sunset at Trebarwith Strand with incoming tide, North Cornwall, UK. March 2014. *NOT AVAILABLE FOR UK 2016 RETAIL CALENDARS
Juvenile Green turtle (Chelonia mydas) swimming near the surface, split level view, Fakarava atoll lagoon, Tuamotu Archipelago, French Polynesia, Pacific Ocean
Oak tree (Quercus robur) in snow, Hampstead Heath, London, UK, January 2013
Low angle view of trees in the Hoh Rainforest, Hall of Mosses Trail, Olympic National Park, Jefferson County, Washington, USA, June
Honeybee (Apis melifera) visiting Lavender (Lavendula angustifolia) in lavender fields, Valensole Plateau, Alpes Haute Provence, France, July
White waterlily (Nymphaea alba) in Naardermeer bog lake, Holland. May
Loggerhead sea turtle (Caretta caretta) emerging from shell, Dalyan delta, Turkey, July
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
St Benets Level Drainage Mill at sunset. Norfolk Broads, England, UK, February
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Matterhorn (4, 478m) at sunrise, with reflection in Lake Stellisee, Zermatt, Switzerland, September 2012
Common cotton grass (Eriophorum angustifolium) Klein Schietveld, Brasschaat, Belgium, May
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Yellow Anaconda (Eunectes notaeus) in vegetation at the edge of the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Wild cat (Felis sylvestris) in snow, Vosges, France, February
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
Erect-crested penguins (Eudyptes sclateri) feeding young chick. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Erect-crested penguins (Eudyptes sclateri) pair. Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Mining bee / Little flower bee (Anthophora bimaculata) taking nectar and pollen from Common fleabane (Pulicaria dysenterica) flower, with pollen covered scopa, Surrey, England, UK. August
Honey bee (Apis mellifera) on Fleabane (Erigion eriginousus) Newport Marshes Reserve, Gwent, Wales, UK. August
River Brue, at dawn with Glastonbury Tor in background, Somerset, UK, August
North American River Otter (Lontra canadensis) two swimming near the surface with autumn leaves reflected in water, Acadia National Park, Maine, USA, October
Svalbard reindeer (Rangifer tarandus platyrhynchus) with bloody antlers shedding velvet, Spitsbergen, Svalbard, Norway, September
Aerial view of Amazon River, with settlements and secondary rainforest, near Iquitos, Peru. July 2015
African lion (Panthera leo), resting on a rock, surrounded by flies, Masai Mara Game Reserve, Kenya. November
Female Jaguar (Panthera onca palustris) with cub (estimated age 5 months), resting on a fallen tree over the Cuiaba River. Porto Jofre, Northern Pantanal, Mato Grosso State, Brazil, South America
Snares island crested penguin (Eudyptes robustus) in forest, Snares Island, New Zealand
Erect-crested penguin (Eudyptes sclateri) pair in greeting display. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Bay Icelandic horse trotting in the snow, Snaefellsnes Peninsula, Iceland, March
Whooper Swan (Cygnus cygnus) standing in snowfall, Hokkaido, Japan, February
Bengal tiger (Panthera tigris tigris), female resting and looking up Kanha National Park, India
Platypus (Ornithorhynchus anatinus) swimming in lake, Tasmania, Australia
Black-footed rock wallaby (Petrogale lateralis), Cape range National Park, Exmouth, Western Australia
Eastern Gray Squirrel (Sciurus carolinensis) feeding in snow, Acadia National Park, Maine, USA, February
American Buffalo or Bison (Bison bison) group with calves, Yellowstone National Park, Wyoming, USA, May
Royal terns (Sterna maxima) pair crossing bills during courtship behaviour, Fort DeSoto Park, Florida, USA
Horned puffin (Fratercula corniculata) close-up portrait, St. Paul Island, Pribilofs, Alaska, USA, July
Playful tabby male kitten, age 10 weeks
Rural view of rolling countryside with grazing cattle, near Frome, Somerset, UK. July 2014
Harvest mouse (Micromys minutus) on wheat stem, Devon, UK (captive). May
Basalt columns at Vik, the southernmost village in Iceland. February 2014
White tailed eagle (Haliaeetus albicilla) in flight with snowy background, Rausu, Japan, February
European Cave Spider (Meta menardi) in limestone cave. Plitvice Lakes National Park, Croatia. January
Stormy light over fields near Longnor, Peak District National Park, Staffordshire, May 2014