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Stormy light over fields near Longnor, Peak District National Park, Staffordshire, May 2014
Mountain hare (Lepus timidus) in white winter coat stretching - in snowy habitat, Scotland. Highly Commended in the Mammals category of the GDT European Photographer of the Year Competition 2015
Water vole (Arvicola amphibius) feeding at edge of water, Kent, UK, December
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Robin (Erithacus rubecula) singing, Norfolk, England, UK, February
Great white egret (Ardea alba) silhouetted at sunset, Lake Csaj, Pusztaszer, Hungary, February. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Ruppells weaver (Ploceus galbula) male building nest, Oman, May
Kelp gull (Larus dominicanus) on rocky shore with crashing waves at sunset, Bird Island, Lamberts Bay, Western Cape province, South Africa, September 2012
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Masai giraffes (Giraffa camelopardalis tippelskirchi), at sunrise, Masai-Mara Game Reserve, Kenya. March
Black-striped capuchin (Sapajus libidinosus) using rock as a tool to break open palm nut, Parnaiba Headwaters National Park, Piaui, Brazil. August
Dawn over the mountains and Nam Song River at Vang Vieng, Laos, March 2009
Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April
Scarlet macaw (Ara macao) flying, blurred motion. Costa Rica
Black Grouse (Tetrao tetrix) displaying with breath vapor at dawn, Utajarvi, Finland, April
Cheetah (Acinonyx jubatus) female standing below dark storm clouds, Masai-Mara game reserve, Kenya
Nile crocodile (Crocodylus niloticus) making its way through sand to the Rufiji River, Selous Game Reserve, Tanzania
Green turtle (Chelonia mydas) from underneath swimming through the surface, Mayotte Island, Comores, Indian Ocean
Snares-crested penguin (Eudyptes robustus) walking along rocks, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Kimmeridge Bay ledges at sunset, Dorset, UK. September 2014
Koala (Phascolarctos cinereus) mother with joey aged eight months, Queensland, Australia, captive
Cornfield annual summer wildflowers growing on one of the plant charity Landlife sCornfield annual summer wildflowers growing on one of the plant charity Landlifes National Wildflower Farms, St Helens, Merseyside, UK, June
Japanese macaque (Macaca fuscata fuscata) mother with rare white furred baby, Jigokudani Valley, Nagano Prefecture, Japan. June
Baby ground squirrel (Xerus inauris) using its tail for shade, Kgalagadi Transfrontier Park, South Africa. Non-ex
Male Bengal tiger (Panthera tigris tigris) known as Sundar (B2) patrolling territory, Bandhavgarh National Park, Madhya Pradesh, India. Endangered species
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Matterhorn (4, 478m) at sunrise, with reflection in Lake Stellisee, Zermatt, Switzerland, September 2012
Clouds over Lake Baikal, Siberia, Russia, October 2010
Dalmatian pelicans (Pelecanus crispus) fighting for fish. Lake Kerkini, Greece. February. Vulnerable species
European beech forest (Fagus sylvatica) in autumn, view from below in the mist, Ilirska Bistrica, Green Karst, Slovenia, October
Snares Crested penguin (Eudyptes robustus) bathing in freshwater pool, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Espanola mockingbird (Mimus macdonaldi) on beach, Galapagos
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Duke of Burgundy butterfly (Hamearis lucina), Wiltshire, UK, June
Comma butterfly (Polygonia c-album) feeding on wild aster, Sutcliffe Park Nature Reserve, Eltham, London, UK, September
Aquatic warbler (Acrocephalus paludicola) Nemunas River Delta, Lithuania. May. Vulnerable species species
Atlantic Walrus (Odobenus rosmarus) at surface, Svalbard, Norway, Arctic, September
Polar bear (Ursus maritimus) cub at entrance to den, Manitoba, Canada. March
Brown hare (Lepus europaeus) three adults in snow covered field during a snow fall, Derbyshire, UK, February
Roe deer (Capreolus capreolus) stag peering through vegetation, Peerdsbos, Brasschaat, Belgium, July
Reflection in fen on surface of water, Klein Schietveld, Brasschaat, Belgium, November
Demoiselle cranes (Anthropoides virgo) flock, India
Brown Bear (Ursus arctos) mother and 3-4 month cubs foraging for clams. Katmai National Park, Alaska, USA, June
Red fox (Vulpes vulpes) in snow, Yellowstone National Park, Wyoming, USA. Wyoming, USA, January
Great egret (Ardea alba) in flight, Florida, USA, March
Ice on black volcanic sand beach with waves, Jokulsarlon, Southeast Iceland, February
Trevose Lighthouse at dusk, long exposure at high tide, Trevose, Cornwall, UK. January 2015
Whooper swan (Cygnus cygnus) in flight in snowfall, Lake Kussharo, Japan, February