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Red deer (Cervus elaphus) stag in the snow, Scotland, March
Polar bear (Ursus maritimus) young bear rolling around in the snow, on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Grizzly bear (Ursus arctos horribilis) female with cub riding on back, Katmai National Park, Alaska, USA, August
Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos
Red fox (Vulpes vulpes) cubs playing, Vosges, France, May
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Corfe Castle, early morning llight and mist, Corfe Castle, Dorset, UK. September 2014
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Great crested grebe (Podiceps cristatus) carrying chick on back, The Netherlands, May
Plains zebra (Equus quagga) grooming foal, Kruger National Park, South Africa, July
African elephant (Loxodonta africana) herd walking in the plains during the dry season, Masai-Mara Game Reserve, Kenya. October
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Kakapo (Strigops habroptilus) close up showing sensory facial feathers, Codfish Island / Whenua Hou, Southland, New Zealand, February. Critically endangered
Aurora Borealis (Northern Lights) over the Klondike River, Yukon Territories, Canada, September 2013
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Yellow Anaconda (Eunectes notaeus) in vegetation at the edge of the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Gayal / Mithun (Bos frontalis) on river bank. Arunachal Pradesh, North East India. November 2014
Galapagos sea lion (Zalophus wollebaeki) juvenile on beach, Galapagos
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
French Bulldog sitting looking up, close up of muzzle
Bengal male cat stalking
Moss covered fallen trees in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
Roe deer (Capreolus capreolus) drinking, Pusztaszer protected landscape, Kiskunsagi, Hungary, May
Stove-pipe sponge (Aplysina archeri) and Orange elephant ear sponge (Agelas clathrodes) Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean
White lipped peccary (Tayassu pecari) on forest floor, Lago Preto Conservation Concession, Yavari River, Peru
Oleander Hawkmoth (Daphnis nerii) resting, Southern Sicily, Italy. March
Female Jaguar (Panthera onca palustris) with cub (estimated age 5 months), resting on a fallen tree over the Cuiaba River. Porto Jofre, Northern Pantanal, Mato Grosso State, Brazil, South America
Polar bear (Ursus maritimus) juvenile crossing over newly forming pack ice, during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Erect-crested penguin (Eudyptes sclateri) pair in greeting display. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Portrait of Skewbald Icelandic horse in the snow, Snaefellsnes Peninsula, Iceland, March
Svalbard reindeer (Rangifer tarandus platyrhynchus) mother and calves, Svalbard, Norway, February
Common cotton grass (Eriophorum angustifolium) Klein Schietveld, Brasschaat, Belgium, May
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Wild cat (Felis sylvestris) in snow, Vosges, France, February
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
Erect-crested penguins (Eudyptes sclateri) feeding young chick. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Erect-crested penguins (Eudyptes sclateri) pair. Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Espanola mockingbird (Mimus macdonaldi) on beach, Galapagos
Mining bee / Little flower bee (Anthophora bimaculata) taking nectar and pollen from Common fleabane (Pulicaria dysenterica) flower, with pollen covered scopa, Surrey, England, UK. August
Honey bee (Apis mellifera) on Fleabane (Erigion eriginousus) Newport Marshes Reserve, Gwent, Wales, UK. August
River Brue, at dawn with Glastonbury Tor in background, Somerset, UK, August
North American River Otter (Lontra canadensis) two swimming near the surface with autumn leaves reflected in water, Acadia National Park, Maine, USA, October
Comma butterfly (Polygonia c-album) feeding on wild aster, Sutcliffe Park Nature Reserve, Eltham, London, UK, September
Svalbard reindeer (Rangifer tarandus platyrhynchus) with bloody antlers shedding velvet, Spitsbergen, Svalbard, Norway, September
Aerial view of Amazon River, with settlements and secondary rainforest, near Iquitos, Peru. July 2015
African lion (Panthera leo), resting on a rock, surrounded by flies, Masai Mara Game Reserve, Kenya. November
Snares island crested penguin (Eudyptes robustus) in forest, Snares Island, New Zealand
Bay Icelandic horse trotting in the snow, Snaefellsnes Peninsula, Iceland, March
Whooper Swan (Cygnus cygnus) standing in snowfall, Hokkaido, Japan, February
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
St Benets Level Drainage Mill at sunset. Norfolk Broads, England, UK, February
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Matterhorn (4, 478m) at sunrise, with reflection in Lake Stellisee, Zermatt, Switzerland, September 2012
Clouds over Lake Baikal, Siberia, Russia, October 2010
Dalmatian pelicans (Pelecanus crispus) fighting for fish. Lake Kerkini, Greece. February. Vulnerable species
European beech forest (Fagus sylvatica) in autumn, view from below in the mist, Ilirska Bistrica, Green Karst, Slovenia, October
Snares Crested penguin (Eudyptes robustus) bathing in freshwater pool, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Duke of Burgundy butterfly (Hamearis lucina), Wiltshire, UK, June
Aquatic warbler (Acrocephalus paludicola) Nemunas River Delta, Lithuania. May. Vulnerable species species
Atlantic Walrus (Odobenus rosmarus) at surface, Svalbard, Norway, Arctic, September
Polar bear (Ursus maritimus) cub at entrance to den, Manitoba, Canada. March
Brown hare (Lepus europaeus) three adults in snow covered field during a snow fall, Derbyshire, UK, February
Illuminated netdevil (Linophryne arborifera), female specimen, from deepsea Atlantic Ocean near Cape Verde at a depth of 1900m
Aggregation of thousands of Spider crabs (Leptomithrax gaimardii) for moulting, South Australia Basin, Australia. Pacific Ocean
Black-footed rock wallaby (Petrogale lateralis), Cape range National Park, Exmouth, Western Australia
Pronghorn (Antilocapra americana) male, Yellowstone National Park, Wyoming, USA, June
American Buffalo or Bison (Bison bison) group with calves, Yellowstone National Park, Wyoming, USA, May
Group of California sea lions (Zalophus californianus) in a kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Canada goose (Branta canadensis) adult swimming leading five goslings, Ithaca, New York, USA
Chilean flamingos (Phoenicopterus chilensis) group of three in flight, Lago Amarga, Torres del Paine National Park, Patagonia, Chile
Rural view of rolling countryside with grazing cattle, near Frome, Somerset, UK. July 2014
Blakistons Fish Owl (Ketupa blakistoni) perched, Hokkaido, Japan, February
Water vole (Arvicola amphibius) feeding at edge of water, Kent, UK, December
Red squirrel (Sciurus vulgaris) feeding, Black Isle, Scotland, UK, February
Brown hare (Lepus europaeus) adult running in wintery landscape. Derbyshire, UK, January
Otter (Lutra lutra) female grooming in seaweed, Mull, Scotland, England, UK, September
Gannet (Sula bassana) diving, July UK Shetland
American goldfinch (Carduelis tristis) male in breeding plumage, perched in Eastern redbud flowers in spring, New York, USA May
Marine iguana (Amblyrhynchus cristatus) underwater. Fernandina Island. Galapagos, Endemic Species