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African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa
Line of Leaf-cutter ants (Atta sp) carrying leaves, digital composite
King penguins (Aptenodytes patagonicus) at sunrise, Falklands. Highly honoured in the Ocean View Category of the Nature's Best Windland Smith Rice Ocean View Competition 2017
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Water lilies (Nymphaeaceae) seen from below, Cenote Car Wash / Aktun Ha, Mexico. Yucatan peninsula
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Darwins Orchid (Angraecum sesquipedale) species which is pollinated by a long-tongued moth, from Ambila, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, August 2009
Welsh Terrier, bitch with puppies aged 8 weeks in snowy scene
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Chlorophyte or green alga (Volvox aureus) in pond water. UK. The larger spheres are approximately 0.5mm in diameter. Digital composite image
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Three Sperm whales (Physeter macrocephalus) socialising under the surface, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species
Fallow deer (Cervus dama) stags at sunset, Holkham Park, North Norfolk, UK, January
Green turtles (Chelonia mydas) swimming with angelfish, Punta Vicente Roca, Isabela Island, Galapagos
Kiang (Equus kiang) herd grazing on steppe in evening light, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) crossing the Mara River. Northern Serengeti, Serengeti National Park, Tanzania
Sanderling (Calidris alba) flock foraging at sunset, Britanny, France, October
Scarlet elf cup fungi (Sarcoscypha coccinea), Uplyme, Devon, England, UK
White ibis (Eudocimus albus) group of four in flight above clouds, Fort Myers Beach, Gulf Coast, Florida, USA, March
RF- Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Hoatzins (Opisthocomus hoazin) perched in tropical rainforest, Tambopata Reserve, Peru, South America
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Three Red-crowned cranes (Grus japonensis) in flight. Hokkaido, Japan. March
Barbary macaque (Macaca sylvanus), two adults with baby, overlooking town and ships in sea at sunset. Gibraltar Nature Reserve, Gibraltar. August 2018
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues
Aerial view of Reindeer (Rangifer tarandus) fording water whilst being herded by Nenet herders. Nenets Autonomous Okrug, Arctic, Russia, July 2017
Northern gannets (Morus bassanus) in flight in Force 8 gales above raging seas. Shetland Islands, Scotland, UK, September
Red-and-Green Macaws (Ara chloropterus), feeding at the wall of a riverside clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Cluster of Fly agaric mushrooms / fungi (Amanita muscaria) growing in coniferous woodland near Inverness, Scottish Highlands. Scotland, UK, October
Accumulation of land snails (Euryptyxis labiosa), escaping the desert heat by resting on a shrubs bark, Ayn Sahnawt, Sultanate of Oman, February
Madagascar green-emerald giant pill millipedes (Zoosphaerium neptunus) Andasibe-Mantadia National Park. Madagascar
Indian lotus (Nelumbo nucifera) flowers, Melbourne Botanic garden, Victoria, Australia
Schools of Pacific creolefish (Cephalopholis colonus) and Panamic sergeant major (Abudefduf troschelii) swimming through sun beams inside an underwater cavern
Mother of thousands (Kalanchoe daigremontiana), viviparous growth with vegetative propagation of plantlets on leaf margins. In cultivation
Focus stacked, inverted image of Diatoms on a microscope slide. Diatoms are single-celled algae which produce approximately 25% of the oxygen we breathe as well as being responsible for approximately
Aggregation of thousands of moulting Spider crabs (Leptomithrax gaimardii), South Australia Basin, Australia. Pacific Ocean
Three Eurasian lynx (Lynx lynx) kittens, aged six weeks, with their mother. Breeding and reintroduction program, Germany. Captive. July
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) shoal swimming over Turtlegrass (Halophila stipulacea) seagrass bed. Marsa Alam, Egypt
Southern elephant seal (Mirounga leonina) beach master / bull roaring amongst females and pups, in breeding colony. St Andrews Bay, South Georgia. October
Walrus (Odobenus rosmarus) amongst group hauled out on shore. Spitsbergen, Svalbard, Norway. July
Percheron horse, group running uphill through snow, one dappled grey leading black horses, snow covered forest in background. Alberta, Canada. February
White-cheeked gibbon (Nomascus leucogenys) female and two offspring, a male and female, sitting in tree. White-cheeked gibbons are sexually dimorphic with females having golden fur
Yellowfin tuna (Thunnus albacares) hunting mackerel (Scomber colias) Tenerife, Canary Islands